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Bagaraatan

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Extinct genus of theropod dinosaurs

Bagaraatan
Temporal range: Late Cretaceous, 70 Ma PreꞒ O S D C P T J K Pg N
Lower jaw bones
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Pantyrannosauria
Genus: Bagaraatan
Osmólska, 1996
Type species
Bagaraatan ostromi
Osmólska, 1996

Bagaraatan (/'ba-ɣa-raa-tan/ meaning 'small' baɣa + 'carnivorous animal, beast of prey' araatan in Mongolian) is a genus of theropod dinosaur from the Late Cretaceous period. Its fossils were found in the Nemegt Formation of Mongolia. Bagaraatan may have been around 3 to 4 metres (9.8 to 13 ft) in length.

History

1996 skeletal diagram showing holotype and assigned elements that are now known to belong to a different species
Partial mandible

The type species, B. ostromi, was described by Halszka Osmólska in 1996. Initially, the post-cranial (ZPAL MgD-I/108) skeleton had been described as "bird-like", while the skull was noted to exhibit features of several different theropod groups.

The material that warranted this conclusion was later found to be a chimaera of two non-avian dinosaurs, with some of the post-crania (hand bones, left femur, tibiotarsus, and rib) being referred to an indeterminate caenagnathid, possibly Elmisaurus. The material that is considered the holotype, which includes the mandible, axial skeleton, pelvis, and one pedal phalanx, likely indicates that Bagaraatan is an indeterminate tyrannosaurid. Similarities between the material and young specimens of Tyrannosaurus and Tarbosaurus indicate that the holotype represents a juvenile tyrannosaurid, one of the smallest currently known.

Classification

Size compared to a human

Holtz classified Bagaraatan as a basal tyrannosauroid, Coria identified it as a troodontid, and Rauhut placed it in Maniraptora. Mark Loewen et al. placed it in basal Tyrannosauroidea, agreeing with the placement by Holtz.

Below is the cladogram by Loewen et al. in 2013.

Tyrannosauroidea
Proceratosauridae

Proceratosaurus bradleyi

Kileskus aristotocus

Guanlong wucaii

Sinotyrannus kazuoensis

Juratyrant langhami

Stokesosaurus clevelandi

Dilong paradoxus

Eotyrannus lengi

Bagaraatan ostromi

Raptorex kriegsteini

Dryptosaurus aquilunguis

Alectrosaurus olseni

Xiongguanlong baimoensis

Appalachiosaurus montgomeriensis

Alioramus altai

Alioramus remotus

Tyrannosauridae

In their 2024 reassessment of Bagaraatan, Słowiak, Brusatte & Szczygielski determined that the initial material referred to Bagaraatan is chimaeric. They suggested that the material recognized as the holotype can more confidently be identified as a member of the Tyrannosauridae or a closely related tyrannosauroid. When tested in a phylogenetic analysis, Bagaraatan was recovered in a polytomy with derived tyrannosauroids within the Tyrannosauridae. However, they caution that its exact phylogenetic position remains uncertain since juvenile specimens tend to be recovered in more basal positions than adults of the same species. Their results are shown in the cladogram below:

Sinotyrannus

Yutyrannus

Proceratosaurus

Guanlong
Kileskus

Suskityrannus

Timurlengia
Xiongguanlong

Dryptosaurus

Raptorex

Tyrannosauridae

Bagaraatan

Bistahieversor

Qianzhousaurus
Alioramus altai
Alioramus remotus
Appalachiosaurus
Albertosaurinae

Tyrannosaurinae

References

  1. ^ Słowiak, Justyna; Brusatte, Stephen L; Szczygielski, Tomasz (2024-02-16). "Reassessment of the enigmatic Late Cretaceous theropod dinosaur, Bagaraatan ostromi". Zoological Journal of the Linnean Society. 202 (3). doi:10.1093/zoolinnean/zlad169. ISSN 0024-4082.
  2. O. W. M. Rauhut (2003). The interrelationships and evolution of basal theropod dinosaurs. Special Papers in Palaeontology 69: 1-213.
  3. ^ Loewen, M.A.; Irmis, R.B.; Sertich, J.J.W.; Currie, P. J.; Sampson, S. D. (2013). Evans, David C (ed.). "Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans". PLoS ONE. 8 (11): e79420. Bibcode:2013PLoSO...879420L. doi:10.1371/journal.pone.0079420. PMC 3819173. PMID 24223179.

Sources

  • Osmolska, H. (1996). "An unusual theropod dinosaur from the Late Cretaceous Nemegt Formation of Mongolia". Acta Palaeontologica Polonica 41; 1-38

External links

[REDACTED] Media related to Bagaraatan at Wikimedia Commons [REDACTED] Data related to Bagaraatan at Wikispecies

Theropoda
Avemetatarsalia
Theropoda
    • see below↓
Theropoda
Theropoda
Neotheropoda
Coelophysoidea
Coelophysidae
Averostra
    • see below↓
Dubious neotheropods
Coelophysis bauri Dilophosaurus wetherilli
Averostra
Averostra
Ceratosauria
Ceratosauridae
Abelisauroidea
Noasauridae
Elaphrosaurinae
Noasaurinae
Abelisauridae
Majungasaurinae
Carnotaurinae
Brachyrostra
Furileusauria
Tetanurae
    • see below↓
Ceratosaurus nasicornis Limusaurus inextricabilis Rajasaurus narmadensis Aucasaurus garridoi
Tetanurae
Tetanurae
Orionides
Megalosauroidea
Piatnitzkysauridae
Megalosauria
Megalosauridae
Megalosaurinae
Afrovenatorinae
Spinosauridae
Baryonychinae
Ceratosuchopsini
Spinosaurinae
Spinosaurini
Avetheropoda
    • see below↓
Piatnitzkysaurus floresi

Torvosaurus tanneri

Spinosaurus aegyptiacus
Avetheropoda
Avetheropoda
Carnosauria
Allosauroidea
Metriacanthosauridae
Metriacanthosaurinae
Allosauria
Allosauridae
Carcharodontosauria
Neovenatoridae
Carcharodontosauridae
Carcharodontosaurinae
Giganotosaurini
Megaraptora?
Megaraptoridae
Coelurosauria
    • see below↓
Xuanhanosaurus qilixiaensis Allosaurus fragilis

Neovenator salerii Carcharodontosaurus saharicus

Australovenator wintonensis
Coelurosauria
Coelurosauria
Tyrannoraptora
Tyrannosauroidea
Coeluridae?
Proceratosauridae
Pantyrannosauria
Eutyrannosauria
Tyrannosauridae
Albertosaurinae
Tyrannosaurinae
Alioramini
Daspletosaurini
Teratophoneini
Tyrannosaurini
Maniraptoromorpha
    • see below↓
Dubious coelurosaurs
Zuolong salleei Stokesosaurus clevelandi

Alioramus remotus

Tarbosaurus bataar
Maniraptoromorpha
Maniraptoromorpha
Neocoelurosauria
Compsognathidae
Maniraptoriformes
Ornithomimosauria
Macrocheiriformes
Deinocheiridae
Ornithomimidae
Maniraptora
Sinosauropteryx prima

Deinocheirus mirificus

Qiupalong henanensis
Taxon identifiers
Bagaraatan
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