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Clavelina
Clavelina picta
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Subphylum:	Tunicata
Class:	Ascidiacea
Order:	Aplousobranchia
Family:	Clavelinidae
Genus:	Clavelina Savigny, 1816

Clavelina ("little bottle") is genus of sea squirts (the Ascidiacea), containing the following species:

• Clavelina amplexa Kott, 2002
• Clavelina arafurensis Tokioka, 1952
• Clavelina auracea Monniot, 1997
• Clavelina australis (Herdman, 1899)
• Clavelina baudinensis Kott, 1957
• Clavelina borealis Savigny, 1816
• Clavelina brasiliensis (Millar, 1977)
• Clavelina breve Monniot, 1997
• Clavelina coerulea Oka, 1934
• Clavelina concrescens Hartmeyer, 1924
• Clavelina cyclus Tokioka & Nishikawa, 1975
• Clavelina cylindrica (Quoy & Gaimard, 1834)
• Clavelina dagysa (Kott, 1957)
• Clavelina dellavallei (Zirpolo, 1825)
• Clavelina detorta (Sluiter, 1904)
• Clavelina elegans (Oka, 1927)
• Clavelina enormis Herdman, 1880
• Clavelina fasciculata Van Name, 1945
• Clavelina fecunda (Sluiter, 1904)
• Clavelina gemmae Turon, 2005
• Clavelina huntsmani Van Name, 1931
• Clavelina kottae (Millar, 1960)
• Clavelina lepadiformis (Müller, 1776)
• Clavelina maculata Monniot & Monniot, 2001
• Clavelina meridionalis (Herdman, 1891)
• Clavelina michaelseni Millar, 1982
• Clavelina miniata Watanabe & Tokioka, 1973
• Clavelina minuta Tokioka, 1962
• Clavelina mirabilis Kott, 1972
• Clavelina moluccensis (Sluiter, 1904)
• Clavelina neapolitana Della Valle, 1881
• Clavelina nigra Kott, 1990
• Clavelina obesa Nishikawa & Tokioka, 1976
• Clavelina oblonga Herdman, 1880
• Clavelina oliva Kott, 1990
• Clavelina ossipandae Hasegawa & Kahijira, 2024
• Clavelina ostrearium (Michaelsen, 1930)
• Clavelina phlegraea Salfi, 1929
• Clavelina picta (Verrill, 1900)
• Clavelina polycitorella (Tokioka, 1954)
• Clavelina pseudobaudinensis (Kott, 1976)
• Clavelina puertosecensis Millar & Goodbody, 1974
• Clavelina robusta Kott, 1990
• Clavelina roseola Millar, 1955
• Clavelina sabbadini Brunetti, 1987
• Clavelina simplex Kott, 2006
• Clavelina steenbrasensis Millar, 1955
• Clavelina viola Tokioka & Nishikawa, 1976

Phylogeny

A 2024 study of cytochrome c oxidase subunit I gene sequences found that Clavelina was paraphyletic to the genus Nephtheis.

References

1. Savigny, 1816. Mémoires sur les Animaux sans Vertèbres, &c. Paris, 1816
2. Karen Sanamyan & Claude Monniot (2011). Noa Shenkar; Arjan Gittenberger; Gretchen Lambert; Marc Rius; Rosana Moreira Da Rocha & Billie J. Swalla (eds.). "Clavelina". Ascidiacea World Database. World Register of Marine Species. Retrieved June 25, 2011.
3. Hasegawa, Naohiro; Kajihara, Hiroshi (2024). "Graveyards of Giant Pandas at the Bottom of the Sea? A Strange-Looking New Species of Colonial Ascidians in the Genus Clavelina (Tunicata: Ascidiacea)". Species Diversity: 53–64. doi:10.12782/specdiv.29.53. Retrieved 19 February 2024.

v t e Tunicate genera
Kingdom: Animalia Phylum: Chordata Clade: Olfactores
Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia Fritillaria Tectillaria Oikopleuridae Althoffia Bathochordaeus Chunopleura Folia Megalocercus Mesochordaeus Mesoikopleura Oikopleura Pelagopleura Sinisteroffia Stegosoma Stolidobranchia (Pleurogona)¹ Molgulidae Anomopera Asajirus Bostrichobranchus Eugyra Fungulus Gamaster Minipera Molgula Molguloides Namiella Oligotrema Paramolgula Pareugyrioides Protomolgula Rhizomolgula Pyuridae Bathypera Bathypyura Boltenia Bolteniopsis Claudenus Cratostigma Ctenyura Culeolus Halocynthia Hartmeyeria Hemistyela Herdmania Heterostigma Microcosmus Paraculeolus Pyura Styelidae Alloeocarpa Amphicarpa Arnbackia Asterocarpa Bathyoncus Bathystyeloides Berillia Botryllocarpa Botrylloides Botryllus Chorizocarpa Cnemidocarpa Dendrodoa Dextrocarpa Dicarpa Distomus Eusynstyela Gynandrocarpa Kukenthalia Metandrocarpa Monandrocarpa Oculinaria Oligocarpa Pelonaia Podostyela Polyandrocarpa Polycarpa Polyzoa Protostyela Psammostyela Seriocarpa Stolonica Styela Symplegma Syncarpa Theodorella Tibitin Thaliacea Pyrosomatidae Pyrosoma Pyrosomella Pyrostremma Salpidae Brooksia Cyclosalpa Helicosalpa Iasis Ihlea Metcalfina Pegea Ritteriella Salpa Soestia Thalia Thetys Traustedtia Doliolida Doliopsoides Paradoliopsis Pseudusa Doliopsidae Doliolula Doliopsis Doliolidae Dolioletta Doliolina Dolioloides Doliolum Enterogona¹ see below↓
Enterogona Aplousobranchia Euherdmania Placentela Stomozoa Vitrum Clavelinidae Clavelina Euclavella Nephtheis Pycnoclavella Diazonidae Diazona Pseudodiazona Pseudorhopalaea Rhopalaea Tylobranchion Didemnidae Atriolum Clitella Coelocormus Didemnum Diplosoma Leptoclinides Lissoclinum Polysyncraton Trididemnum Holozoidae Distaplia Hypodistoma Hypsistozoa Neodistoma Polydistoma Protoholozoa Pseudoplacentela Scotiazoa Sigillina Sycozoa Polycitoridae Archidistoma Brevicollus Cystodytes Eucoelium Eudistoma "Exostoma" Millarus Polycitor Polycitorella Rhombifera Salix Polyclinidae Aplidiopsis Aplidium Morchellioides Morchellium Neodictyon Polyclinella Polyclinum Sidneioides Synoicum Protopolyclinidae Condominium Monniotus Protopolyclinum Pseudodistomidae Anadistoma Citorclinum Pseudodistoma Ritterellidae Dumus Pharyngodictyon Ritterella Phlebobranchia Dimeatus Hypobythius Agneziidae Adagnesia Agnezia Caenagnesia Proagnesia Pterygascidia Ascidiidae Ascidia Ascidiella Fimbrora Phallusia Psammascidia Cionidae Araneum Ciona Tantillulum Corellidae Abyssascidia Chelyosoma Clatripes Corella Corelloides Corellopsis Corynascidia Dextrogaster Mysterascidia Rhodosoma Xenobranchion Octacnemidae Benthascidia Cibacapsa Cryptia Dicopia Kaikoja Megalodicopia Myopegma Octacnemus Polyoctacnemus Situla Perophoridae Ecteinascidia Perophora Plurellidae Microgastra Plurella
Tunicate-like fossils Ediacaran Ausia Burykhia Finkoella Pharyngomorpha Yarnemia Cambrian Cheungkongella Megasiphon Shankouclava Later Palaeoquadrum Quadrifolium Toscanisoma Khmeriamorpha Khmeria Zardinisoma Ichnotaxa Catellocaula
¹Part of the paraphyletic class Ascidiacea Category [REDACTED] Commons

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