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Nanotitanops Temporal range: Lutetian PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Perissodactyla
Family:	†Brontotheriidae
Genus:	†Nanotitanops Qi & Beard, 1998
Species:	†N. shanghuangensis
Binomial name
†Nanotitanops shanghuangensis Qi & Beard, 1998

Nanotitanops is an extinct genus of Brontothere from the middle Eocene of China. It contains a single species, N. shanghuangensis. It is known only from isolated teeth, the smallest of any known Brontothere.

Description

This genus is currently known only from dental evidence. Described specimens do not exceed 13.55 millimeters in length and 12 millimeters in width. While it retains some basal traits, the teeth possess features which point toward it being relatively derived. It differs from basal designs in having relatively longer and narrower molars with stronger development of buccal shearing crests and only a vestigial paraconule on the upper molars. The dental morphology is most similar to the much larger genera Epimanteoceras and Rhinotitan.

Relationships

The exact placement of Nanotitanops within Brontotheridae is uncertain. There are other small brontotheres from the Eocene of Asia such as Microtitan, Pygmaetitan, and Protitan though morphological evidence does not support a monophyletic relationship between them. Morphological analysis tends to put Nanotitanops in a polytomy near the node of the subfamily Brontotheriinae. It has been proposed as a possible sister taxon to Epimanteoceras formosa. Nanotitanops was originally named Nanotitan, but this genus name was preoccupied by an orthopteroid insects from the Triassic of Kyrgyzstan. The genus name is derived from the Greek nanos, meaning dwarf, titan, referring to the Titans of Greek mythology, and the Greek suffix -ops, meaning face. The specific epithet of the only species refers to the village in which the first and only specimens have been found.

Paleobiology

This genus is only described from the astoundingly rich middle Eocene fissure-filling near the village of Shanghuang in southern Jiangsu Province, China. The fissures also held another small brontothere, Microtitan cf. mongoliensis, a species known from the Irdin Manha Formation of the Nei Mongol Autonomous Region of China. They are both known only from a site dubbed Fissure D. This fissure contained various other mammalian fauna including the primitive cricetid rodent Pappocricetodon antiquus and the omomyid or omomyid-like primate Macrotarsius macrorhysis. The nearby fissures have included a large diversity of primates. There is also a Dichobunid known from the area, called Elaschitotherium qii.

References

1. ^ Qi, Tao; Beard, K. Christopher (1996-09-19). "Nanotitan shanghuangensis, gen. et sp. nov.: the smallest known brontothere (Mammalia: Perissodactyla)". Journal of Vertebrate Paleontology. 16 (3): 578–581. doi:10.1080/02724634.1996.10011342. ISSN 0272-4634.
2. ^ Qi, Tao; Beard, K. Christopher (1998-12-28). "Nanotitanops, a new name for Nanotitan Qi and Beard, 1996, not Nanotitan Sharov, 1968". Journal of Vertebrate Paleontology. 18 (4): 812–812. doi:10.1080/02724634.1998.10011109. ISSN 0272-4634.
3. ^ Li, S. (2018). A new species of Brontotheriidae from the Middle Eocene of Junggar Basin, Xinjiang, China. Vertebrata PalAsiatica, 56(1), 25-44.
4. Mihlbachler, M. C. (2007). Eubrontotherium clarnoensis, a new genus and species of brontothere (Brontotheriidae, Perissodactyla) from the Hancock Quarry, Clarno Formation, Wheeler County, Oregon. PaleoBios, 27(1), 19-39.
5. Mihlbachler, M. C., & Samuels, J. X. (2016). A small-bodied species of Brontotheriidae from the middle Eocene Nut Beds of the Clarno Formation, John Day Basin, Oregon. Journal of Paleontology, 90(6), 1233-1244.
6. Beard, K. C., Qi, T., Dawson, M. R., Wang, B., & Li, C. (1994). A diverse new primate fauna from middle Eocene fissure-fillings in southeastern China. Nature, 368(6472), 604-609.
7. GRÉGOIRE, M., GUO, J., & BEARD, K. C. (2004). A new small dichobunid artiodactyl from Shanghuang (middle Eocene, eastern China): implications for the early evolution of proto-selenodonts in Asia. Bulletin of Carnegie Museum of Natural History, 2004(36), 177-197.

v t e Perissodactyla
Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Perissodactyla
Perissodactyla Incertae sedis †Ghazijhippus †Mesolambdolophus †Olbitherium †Anthracobunia Radinskya †Anthracobunidae Anthracobune Hsanotherium Jozaria Obergfellia †Cambaytheriidae Cambaytherium Kalitherium Nakusia Perissobune †Brontotheriidae See Brontotheriidae Equoidea †Cymbalophus †Hallensia †Pliolophus †Systemodon Equidae See Equidae †Palaeotheriidae †Pachynolophinae Anchilophus Bepitherium Eurohippus Hyracotherium Lijiangia Lophiohippus Lophiotherium Metanchilophus Orolophus Pachynolophus Paranchilophus Propalaeotherium Qianohippus †Palaeotheriinae Cantabrotherium Iberolophus Idiodontherium Leptolophus Mekodontherium Metaplagiolophus Palaeotherium Paraplagiolophus Plagiolophus Pseudopalaeotherium †Hyopsodontidae? Aletodon Chacomylus Dorraletes Haplaletes Haplomylus Hyopsodus Litomylus Louisina Monshyus Paratricuspiodon Paschatherium Phenacodaptes Tricuspiodon †Phenacodontidae? Tetraclaenodon †Meniscotheriinae Ectocion Meniscotherium †Phenacodontinae Almogaver Copecion Eodesmatodon Lophocion Phenacodus Tapiromorpha See below↓
Tapiromorpha Incertae sedis †Cambaylophus †Skopaiolophus †Isectolophidae Chowliia Gandheralophus Karagalax Meridiolophus Orientolophus Sastrilophus †Homogalaxinae Homogalax †Isectolophinae Isectolophus Schizolophodon †Ancylopoda See Ancylopoda Ceratomorpha Tapiroidea †Deperetellidae Bahinolophus Deperetella Irenolophus Teleolophus †Helaletidae Colodon Dilophodon Helaletes Heptodon Heteraletes Paracolodon Plesiocolopirus Thuliadanta? Vastanolophus Tapiridae †Eotapirus †Miotapirus †Nexuotapirus †Paratapirus †Plesiotapirus †Protapirus †Tapiravus Tapirus Rhinocerotoidea See below↓
Rhinocerotoidea Rhinocerotoidea sensu lato †Breviodon †Fouchia †Minchenoletes †Siamolophus †Triplopus †Yimengia †Indolophidae Indolophus Rhinocerotoidea sensu stricto †Selenaletes †Hyrachyidae Hyrachyus †Eggysodontidae Allacerops Eggysodon Guangnanodon Proeggysodon †Hyracodontidae Ardynia Ephyrachyus Epitriplophus Hyracodon Prohyracodon Rhodopagus Triplopides †Amynodontidae Amynodontopsis Armania Cadurcotherium Caenolophus Huananodon Hypsamynodon Procadurcodon Rostriamynodon Teilhardia †Amynodontinae Amynodon †Cadurcodontini Cadurcodon Lushiamynodon Sharamynodon Sianodon †Metamynodontini Gigantamynodon Megalamynodon Metamynodon Paramynodon Zaisanamynodon †Paraceratheriidae †Forstercooperiinae Forstercooperia Gobicerops Pappaceras †Paraceratheriinae Aralotherium Dzungarotherium Juxia Paraceratherium Urtinotherium Rhinocerotidae See below↓
Rhinocerotidae Rhinocerotidae †Amphicaenopus †Epiaceratherium †Guixia †Teletaceras †Uintaceras †Woodoceras †Aceratheriinae Aceratherium Acerorhinus Alicornops Aphelops Chilotheridium Chilotherium Floridaceras Galushaceras Hoploaceratherium Mesaceratherium Peraceras Persiatherium Plesiaceratherium Ronzotherium Shansirhinus Sinorhinus Subchilotherium †Elasmotheriinae Gulfoceras Victoriaceros †Diceratheriini Diceratherium Penetrigonias Skinneroceras Subhyracodon Trigonias †Elasmotheriini Bugtirhinus Caementodon Elasmotherium Hispanotherium Iranotherium Kenyatherium Meninatherium Menoceras Ninxiatherium Ningxiatherium Ougandatherium Parelasmotherium Procoelodonta Sinotherium Tongxinotherium Rhinocerotinae †Dromoceratherium †Lartetotherium †Molassitherium †Protaceratherium †Teleoceratini Aprotodon Brachydiceratherium Brachypotherium Diaceratherium Prosantorhinus Shennongtherium Teleoceras Rhinocerotina Dicerorhinini †Coelodonta Dicerorhinus †Dihoplus †Pliorhinus †Stephanorhinus Dicerotini Ceratotherium Diceros †Miodiceros †Paradiceros Rhinocerotini †Gaindatherium †Nesorhinus Rhinoceros †Rusingaceros

v t e Brontotheriidae
Kingdom: Animalia Phylum: Chordata Clade: Synapsida Class: Mammalia Order: Perissodactyla Suborder: Hippomorpha
Basal brontotheres	Danjiangia Eotitanops Lambdotherium Mulkrajanops Pakotitanops Palaeosyops
Brontotheriinae	Acrotitan Arctotitan Bunobrontops Desmatotitan Dolichorhinus Fossendorhinus Hyotitan Mesatirhinus Metarhinus Metatelmatherium Microtitan Nanotitanops Pygmaetitan Qufutitan Sphenocoelus Sthenodectes Telmatherium Brontotheriini Diplacodon Protitan Protitanotherium Rhinotitan Pachytitan Brachydiastematherium Sivatitanops Embolotheriina Aktautitan Embolotherium Gnathotitan Maobrontops Metatitan Nasamplus Protembolotherium Brontotheriina Diantotitan Duchesneodus Epimanteoceras Eubrontotherium Megacerops Notiotitanops Parabrontops Parvicornus Protitanops

• Eocene animals of Asia
• Brontotheres
• Brontotheres of Asia
• Clarno Formation