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This article records new taxa of fossil archosaurs of every kind that are scheduled described during the year 2022, as well as other significant discoveries and events related to paleontology of archosaurs that are scheduled to occur in the year 2022.

Pseudosuchians

New pseudosuchian taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Confractosuchus	Gen. et sp. nov	In press	White et al.	Late Cretaceous (Cenomanian)	Winton Formation	Australia	A eusuchian. The type species is C. sauroktonos.
Diplocynodon kochi	Sp. nov	Valid	Venczel & Codrea	Eocene (Priabonian)	Cluj Limestone Formation	Romania
Eptalofosuchus	Gen. et sp. nov	Valid	Marinho et al.	Late Cretaceous	Uberaba Formation	Brazil	A notosuchian crocodylomorph. The type species is E. viridi. Announced in 2021; the final article version was published in 2022.
Hanyusuchus	Gen. et sp. nov		Iijima et al.	Holocene		China	A member of the family Gavialidae with a mosaic of gavialine and tomistomine features across the skeleton. The type species is H. sinensis.
Kinyang	Gen. et spp. nov		Brochu et al.	Early - Middle Miocene	Maboko Formation Lokone Formation	Kenya	A broad skulled genus of osteolaemine crocodile. Type species is K. mabokoensis, also includes new species K. tchernovi.
Mambawakale	Gen. et sp. nov	Valid	Butler et al.	Middle Triassic	Manda Beds	Tanzania	An early diverging pseudosuchian of uncertain affinities. The type species is M. ruhuhu.
Maomingosuchus acutirostris	Sp. nov	Valid	Massonne et al.	Eocene (late Bartonian–Priabonian)	Na Duong Formation	Vietnam
Sacacosuchus	Gen. et sp. nov		Salas-Gismondi et al.	Late Miocene	Pisco Formation	Peru	A member of the family Gavialidae. The type species is S. cordovai.
Yanjisuchus	Gen. et sp. nov	Valid	Rummy et al.	Cretaceous (Albian–Cenomanian)	Longjing Formation	China	A paralligatorid crocodyliform. The type species is Y. longshanensis. Announced in 2021; the final article version was published in 2022.

General pseudosuchian research

• A study on the mandible embryogenesis in extant caimans, and on its implications for the knowledge of the evolution of postdentary lower jaw of pseudosuchians, is published by Bona et al. (2022).
• A study on the musculature of jaws of crocodylians and fossil suchians, and on its implications for the knowledge of the impact of skull flattening on muscle anatomy of suchians, is published by Sellers et al. (2022).
• Revision of Tsylmosuchus donensis and Scythosuchus basileus is published by Sennikov (2022), who interprets the latter taxon as a junior synonym of the former one, and interprets T. donensis as a likely member of the family Ctenosauriscidae.
• Partial maxilla of a basal loricatan is described from the Upper Triassic (Carnian) lower Candelária Sequence of the Hyperodapedon Assemblage Zone (Brazil) by Damke et al. (2022), expanding known record of loricatans in this unit.
• A study aiming to model to the likely gait of Batrachotomus kupferzellensis is published by Polet & Hutchinson (2022).
• A study on the feeding ecology of Batrachotomus kupferzellensis is published by Mujal et al. (2022).

Aetosaur research

• A study on the microstructure of the humerus, femur and tibia of Aetosauroides scagliai, and on its implications for the knowledge of the paleobiology of this aetosaur, is published by Ponce, Desojo & Cerda (2022).

Crocodylomorph research

• A study on the evolution of palatal anatomy in early crocodylomorphs is published by Dollman & Choiniere (2022).
• Description of the anatomy of the holotype specimen of Junggarsuchus sloani, its comparison to Dibothrosuchus elaphros, and a study on the affinities of both taxa is published by Ruebenstahl et al. (2022).
• Review of the type material of the crocodylomorph ichnotaxon Crocodylopodus meijidei from the Berriasian of Spain, and a study on the locomotion of the trackmaker, is published by Castanera et al. (2022).
• Description of the microstructure of the tooth and tooth attachment tissues of Notosuchus terrestris is published by Navarro et al. (2022), who find the relative and absolute enamel thickness of N. terrestris to be similar to those reported for carnivorous notosuchians such as baurusuchids, and interpret their findings as indicating that the tooth growth rates of N. terrestris were reduced in comparison with other notosuchians.
• A study on the bone histology of Mariliasuchus amarali, based on a new specimen is published by Sena et al. (2022).
• A study on the palatal anatomy of Sebecus icaeorhinus and its implications for the knowledge of the variability in the palatal anatomy within Sebecidae, is published by Bravo et al. (2022).
• A study on the phylogenetic relationships of neosuchians and on timing of the origination of key clades in neosuchian evolution is published by Groh et al. (2022).
• A study on the evolution of salt glands in thalattosuchians is published by Cowgill et al. (2022).
• A study on the phylogenetic affinities of Portugalosuchus azenhae is published by Darlim et al. (2022).
• Lindblad et al. (2022) describe specimens of Borealosuchus griffithi and B. sternbergii preserved in proximity to each other within the same deposits of the Paleocene Ravenscrag Formation (Saskatchewan, Canada), expanding known stratigraphic and geographic range of these species, and interpret this finding as possible evidence of niche partitioning or other ecological relationships between the two species.
• Reconstructions of the inner cavities of the holotype skulls of Arenysuchus gascabadiolorum and Agaresuchus subjuniperus are presented by Puértolas-Pascual et al. (2022).
• Kuzmin (2022) reviews all known material of long-snouted crocodyliforms from the Upper Cretaceous (Cenomanian–Santonian) of Central Asia, confirms Zholsuchus procerus to be a valid taxon, and argues that it is likely one of the oldest known members of the crown group of Crocodylia.
• Redescription of the holotype of Notocaiman stromeri, and a study on its taxonomic status and phylogenetic affinities, is published by Bona et al. (2022).
• The oldest known fossil material of a member of the genus Crocodylus in Madagascar, dated to between 7670 and 7510 years cal BP, is described by Martin et al. (2022).

Non-avian dinosaurs

New dinosaur taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Abditosaurus	Gen. et sp. nov	Valid	Vila et al.	Late Cretaceous (Maastrichtian)	Conques Formation	Spain	A saltasaurine titanosaur. The type species is A. kuehnei.
Bashanosaurus	Gen. et sp. nov	Valid	Dai et al.	Middle Jurassic (Bajocian)	Shaximiao Formation	China	A basal stegosaur. The type species is B. primitivus.
Dzharaonyx	Gen. et sp. nov	Valid	Averianov & Sues	Late Cretaceous (Turonian)	Bissekty Formation	Uzbekistan	An alvarezsaurid theropod. The type species is D. eski.
Guemesia	Gen. et sp. nov	In press	Agnolín et al.	Late Cretaceous (Campanian)	Los Blanquitos Formation	Argentina	An abelisaurid theropod. The type species is G. ochoai.
Huallasaurus	Gen. et sp. nov	Valid	Rozadilla et al.	Late Cretaceous (Maastrichtian)	Los Alamitos Formation	Argentina	A saurolophine hadrosaurid belonging to the tribe Kritosaurini. The type species is 'Kritosaurus' australis (Bonaparte, 1984).
Iberospinus	Gen. et sp. nov	Valid	Mateus & Estraviz-López	Early Cretaceous (Barremian)	Papo Seco Formation	Portugal	A spinosaurid theropod. The type species is I. natarioi.
Kelumapusaura	Gen. et sp. nov	Valid	Rozadilla et al.	Late Cretaceous (Campanian-Maastrichtian)	Allen Formation	Argentina	A saurolophine hadrosaurid belonging to the tribe Kritosaurini. The type species is K. machi.
Maip	Gen. et sp. nov	Valid	Aranciaga Rolando et al.	Late Cretaceous (Maastrichtian)	Chorrillo Formation	Argentina	A megaraptorid theropod. The type species is M. macrothorax.
Menucocelsior	Gen. et sp. nov	Valid	Rolando et al.	Late Cretaceous (Campanian-Maastrichtian)	Allen Formation	Argentina	A titanosaur sauropod. The type species is M. arriagadai. Announced in 2021; the final article version was published in 2022.
Napaisaurus	Gen. et sp. nov	Valid	Ji & Zhang	Early Cretaceous	Xinlong Formation	China	A basal member of Iguanodontia. The type species is N. guangxiensis. Announced in 2021; the final article version was published in 2022.
Ondogurvel	Gen. et sp. nov	In press	Averianov & Lopatin	Late Cretaceous (Campanian)	Barun Goyot Formation	Mongolia	An alvarezsaurid theropod. The type species is O. alifanovi.
Papiliovenator	Gen. et sp. nov	Valid	Pei et al.	Late Cretaceous (Campanian)	Bayan Mandahu Formation	China	A troodontid theropod. The type species is P. neimengguensis. Announced in 2021; the final article version to be published in 2022.
Paralitherizinosaurus	Gen. et sp. nov	Valid	Kobayashi et al.	Late Cretaceous (Campanian)	Osoushinai Formation	Japan	A therizinosaurid theropod. The type species is P. japonicus.
Sierraceratops	Gen. et sp. nov	Valid	Dalman et al.	Late Cretaceous (latest Campanian–Maastrichtian)	Hall Lake Formation	United States ( New Mexico)	A chasmosaurine ceratopsid. The type species is S. turneri. Announced in 2021; the final article version will be published in 2022.
Tyrannosaurus imperator	Sp. nov	In press	Paul, Persons & Van Raalte	Late Cretaceous (late Maastrichtian)	Hell Creek, Lance, Laramie, Arapahoe, McRae?, North Horn?, and Javelina? Formations	United States ( Montana,  North Dakota,  South Dakota,  Wyoming,  New Mexico?,  Texas?,  Utah?)	A tyrannosaurine; a proposed species of Tyrannosaurus.
Tyrannosaurus regina	Sp. nov	In press	Paul, Persons & Van Raalte	Late Cretaceous (late Maastrichtian)	Hell Creek, Lance, Ferris, Denver, Frenchman, Willow Creek, and Scollard Formations	Canada ( Alberta,  Saskatchewan)  United States ( Colorado,  Montana,  North Dakota,  South Dakota,  Wyoming)	A tyrannosaurine; a proposed species of Tyrannosaurus.
Yuxisaurus	Gen. et sp. nov	Valid	Yao et al.	Early Jurassic (Sinemurian–Toarcian)	Fengjiahe Formation	China	An early thyreophoran. The type species is Y. kopchicki.

General non-avian dinosaur research

• A study on the ecomorphospace occupation of major megaherbivorous dinosaur clades from the Late Jurassic through to the end of the Late Cretaceous in North America is published by Wyenberg-Henzler (2022).
• A study on the age of a new sauropod-dominated dinosaur fauna from the Lower Shaximiao Formation in Yunyang (Chongqing, China) is published by Zhou et al. (2022).
• Description of theropod and ornithischian tracks from the Jurassic Imilchil and Isli formations (Morocco), including theropod tracks representing the ichnogenus Changpeipus (otherwise known from abundant occurrences in East Asia, and possibly indicative of faunal exchange between East Asia and northern Africa in the Middle Jurassic), is published by Klein et al. (2022).
• Revision of the Early Cretaceous dinosaur assemblage from the Mogoito locality (Murtoi Formation, Transbaikalia, Russia) is published by Averianov et al. (2022), who report the presence of Tengrisaurus starkovi, as well as ornithomimosaur, therizinosaur, dromaeosaurid, jeholosaurid (including teeth previously attributed to Psittacosaurus) and indeterminate sauropod and theropod fossil material.
• New fossil site for the Jehol Biota, preserving faunal assemblage dominated by dinosaur fossils and similar in composition to the Lujiatun Unit of the Yixian Formation at Beipiao (Liaoning, China), is reported from Ningcheng (Inner Mongolia, China) by Zhang et al. (2022).
• Two trackways produced by at least two different dinosaur taxa (smaller, bipedal trackmaker, likely a theropod, and a larger trackmaker of uncertain affinities, possibly a theropod or ornithopod), representing the first records of non-avian dinosaurs from Palestine reported to date, are described from a new site located within the city of Al-Bireh and belonging to the Albian Soreq Formation by Lallensack et al. (2022).
• A large dinosaur tracksite preserving theropod tracks and abundant hadrosaurid tracks is described from the Upper Cretaceous (Campanian) Wapiti Formation (Alberta, Canada) by Enriquez et al. (2022), who evaluate the implications of this finding for the knowledge of the paleoecology of dinosaurs known from the Wapiti Formation.
• A study on the calcium isotope variability in tooth enamel of dinosaurs from the Upper Cretaceous Dinosaur Park Formation, Horseshoe Canyon Formation and Scollard Formation (Alberta, Canada), and on its implications for the knowledge of the stability of food web structure of non-avian dinosaur communities in the millions of years preceding the end of the Cretaceous, is published by Martin et al. (2022).
• A study on the stable isotope compositions of dinosaur eggshells and their associated depositional settings of the Upper Cretaceous Huizhou Formation (China), and on their implications for the knowledge of the environment of dinosaur nesting sites, is published by He et al. (2022).
• Review of chemistry of the organic molecules detected in non-avian dinosaur fossils to date is published by Tahoun et al. (2022).

Ornithischian research

• A study on the phylogenetic relationships of the neornithischian dinosaurs commonly referred to as "hypsilophodontids", aiming to determine causes of conflicting placements of these taxa in different phylogenetic analyses, is published by Brown et al. (2022).
• A study aiming to determine whether ornithischian megaherbivores from the upper Oldman Formation (Alberta, Canada) partitioned their niches based on spatial patterns of occupation and resource-use, based on strontium, oxygen and carbon isotope data, is published by Cullen et al. (2022).

Cerapod research

• Description of postcranial material tentatively assigned to Camptosaurus sp. from the Late Jurassic Villar del Arzobispo Formation (Valencia, Spain) is published by Sánchez-Fenollosa et al. (2022)
• Redescription of the holotype of Draconyx loureiroi, including description of previously unreported material, and a study on the phylogenetic affinities of this taxon is published by Rotatori, Moreno-Azanza & Mateus (2022).
• A new specimen of Iguanodon bernissartensis (a partial axial skeleton) is described from the Early Cretaceous (Upper Barremian) Arcillas de Morella Formation (Spain) by Gasulla et al. (2022)
• Description of new fossils of large bodied styracosternans pertaining to two different taxa from the Early Cretaceous El Castellar Formation (Teruel, Spain) is published by García-Cobeña, Verdú,and Cobos (2022), who also describe the first dinosaur tracksite from this formation.
• Fossil material of non-hadrosauriform styracosternans is described from the Lower Cretaceous Khok Kruat Formation by Samathi & Suteethorn (2022), representing the first record of a juvenile iguanodontian co-occurring with an adult (possibly of the same taxon) from Thailand.
• Description of a nearly complete and articulated skeleton of a juvenile hadrosauroid from the Upper Cretaceous Bayan Shireh Formation (Mongolia), distinct from Gobihadros mongoliensis and likely representing a second, previously unknown hadrosauroid taxon from this formation, is published by Averianov, Lopatin & Tsogtbaatar (2022).
• A clutch of subspherical dinosaur eggs, at least two of which contain identifiable hadrosauroid embryos with possible affinities with such taxa as Levnesovia transoxiana, Nanningosaurus dashiensis or Tanius sinensis, is described from the Upper Cretaceous Hekou Formation (China) by Xing et al. (2022).
• Redescription of two putative rhabdodontid braincases from the Maastrichtian of the Haţeg Basin (Romania) is published by Augustin et al. (2022), who reinterpret these specimens as hadrosauroid braincases, likely belonging to members of the species Telmatosaurus transsylvanicus.
• Review of the taxonomic status, phylogenetic relationships and biogeography of hadrosauroids known from Mexico is published by Ramírez-Velasco (2022).
• A study on the morphometric changes within the skull and dietary changes during growth of North American hadrosaurids is published by Wyenberg-Henzler, Patterson & Mallon (2022).
• A pathological ulna of a specimen of Amurosaurus riabinini, preserved with a hypertrophied and swollen distal region and with the distal articular surface engulfed within a large overgrowth of newly formed bone, is described from the Maastrichtian Udurchukan Formation (Amur Region, Russia) by Bertozzo et al. (2022), who interpret the bone as still healing prior to the animal's death, with the misalignment of the fracture and the resulting malunion of the two fragments of the bone probably causing the animal to limp and walk on three limbs.
• A study on the taphonomy of a bonebed with fossils of members of the genus Gryposaurus from the lower unit of the Campanian Oldman Formation (Alberta, Canada), and on the bone microstructure of specimens from this bonebed, is published by Scott et al. (2022).
• Description of the skin of a hadrosaurid specimen (probably belonging to the species Edmontosaurus annectens) from the Maastrichtian Frenchman Formation (Saskatchewan, Canada), preserving unique corrugated scales that have not been observed in this species before, is published by Libke et al. (2022).
• A method which can be used to determine the percent vascularity in any given CT slice of the frontoparietal is presented by Nirody et al. (2022), who use this method to study changes of vascularity in the frontoparietal dome of Stegoceras validum during its ontogeny.
• A study on tooth replacement patterns in Yinlong downsi, Hualianceratops wucaiwanensis and Chaoyangsaurus youngi is published by Hu et al. (2022).
• The oldest umbilical scar reported to date, which is also the first umbilical scar reported to date in a non-avian dinosaur, is described in a specimen of Psittacosaurus from the Lower Cretaceous Jehol Group (China) by Bell et al. (2022).
• A new articulated skeleton of Yamaceratops dorngobiensis, representing the first substantially complete skeleton and the first known juvenile specimen of this taxon, is described from the Upper Cretaceous (?Santonian-Campanian) Javkhlant Formation (Mongolia) by Son et al. (2022).
• A study on the bone histology of Koreaceratops hwaseongensis is published by Baag & Lee (2022).
• Mallon et al.(2022) redescribe two ceratopsid frills from Canada attributed to Torosaurus (representing the northernmost records of this genus reported to date), and evaluate possible implications of these specimens for determination of the status of Torosaurus as a genus distinct from Triceratops.
• A study on the fenestra perforating the right squamosal of the Triceratops horridus specimen known as Big John is published by D'Anastasio et al. (2022), who interpret this fenestra as the result of a traumatic event, possibly a fight with another Triceratops.
• A study on the hadrosaurid and ceratopsid faunas of the Upper Cretaceous Prince Creek Formation, Cantwell Formation and Chignik Formation (Alaska, United States), and on the possible impact of the climate on differences of relative abundances of hadrosaurids and ceratopsids from these formations, is published by Fiorillo et al. (2022).

Thyreophoran research

• Schade et al. (2022) create digital models of the braincase of Struthiosaurus austriacus, and evaluate the implication of its anatomy for the knowledge of the behavioral capacities of this dinosaur.
• Partial skull of a member or a relative of the genus Kunbarrasaurus is described from the Albian Toolebuc Formation by Frauenfelder et al. (2022), representing the oldest ankylosaurian material from Queensland (Australia) reported to date.
• A description of a partial thyreophoran osteoderm from an Early Jurassic Konservatlagerstätte near Grimmen, Germany is published by Schade & Ansorge (2022).

Saurischian research

Sauropodomorph research

• A study on the shape variation of long bones in limbs of sauropodomorphs, and on its implications for the knowledge of the evolution of the sauropod bauplan, is published by Lefebvre et al. (2022).
• Review of the biological mechanisms underpinning the evolutionary transition from obligatory or facultative bipedalism to an obligatory quadrupedalism in sauropodomorphs is published by Otero & Hutchinson (2022).
• Revision and a study on the phylogenetic affinities of Carnian sauropodomorphs from South America is published by Langer et al. (2022).
• A study on the shape and variation of the anterolateral scar in the femora of Pampadromaeus barberenai and Buriolestes schultzi, and on its implications for the knowledge of the distribution of the anterolateral scar in ornithodirans, is published by Müller (2022).
• Reconstruction of the appendicular musculature of Thecodontosaurus antiquus is presented by Ballell, Rayfield & Benton (2022).
• A new, large sized early sauropodomorph specimen is described from the Late Triassic (Carnian) Santa Maria Formation (Brazil) by Müller and Garcia (2022)
• Revision of the non-gravisaurian sauropodiform taxa from South America (Mussaurus patagonicus, Leonerasaurus taquetrensis, Lessemsaurus sauropoides and Ingentia prima is published by Apaldetti & Martínez (2022).
• A study on changes occurring in the postcranial skeleton of Mussaurus patagonicus during its ontogeny is published by Otero & Pol (2022).
• Evidence of widespread incompleteness of necks even in best-preserved and best-known sauropod specimens, and of widespread distortion of known sauropod cervical vertebrae, is presented by Taylor (2022).
• A study aiming to determine whether the sauropod tracks from the Kimmeridgian Courtedoux-Tchâfouè track site (Reuchenette Formation, Switzerland) all represent the same ichnogenus and whether there is variation in their morphology, using linear-based and geometric morphometrics methods, is published by Sciscio et al. (2022).
• Fragmentary heart-shaped tooth crown of a sauropod is described from the Bathonian Jaisalmer Formation (India) by Sharma, Singh & Satheesh (2022), who interpret this specimen as the first known record of a member of Turiasauria from India.
• Description of a nearly complete skull of a member of the genus Apatosaurus from the Upper Jurassic Morrison Formation (Como Bluff, Wyoming, United States), and a study on the tooth replacement in this specimen, is published by Peterson et al. (2022), who interpret their findings as indicative of a different tooth replacement pattern in Apatosaurus relative to Diplodocus, possibly pointing to the ecological niche partitioning among diplodocids and to Apatosaurus’ preference for a food source with tougher vegetation.
• A study on bony pathologic structures stemming from the pneumatic features in the cervical vertebrae of a diplodocine specimen from the Lower O’Hair Quarry (Morrison Formation; Montana, United States) is published by Woodruff et al. (2022), who diagnose this specimen as likely affected by an avian-like airsacculitis, constituting the first identification of this disease in a non-avian dinosaur specimen.
• A study on the external morphology, internal microanatomy and bone microstructure of the hemispinous processes of the vertebrae from the holotype specimen of Amargasaurus cazaui and an indeterminate dicraeosaurid specimen from the La Amarga Formation (Argentina), aiming to reconstruct soft tissues associated with those processes and to determine their functional significance, is published by Cerda, Novas, Carballido and Salgado (2022).
• Four sauropod ribs preserving evidence of three different pathologies (including osteosclerosis) are described from the Middle Jurassic of Yunyang (China) by Tan et al. (2022).
• Revision of the fossil record of non-titanosaur macronarians from South America is published by Carballido, Bellardini & Salgado (2022).
• A study on the morphology, preservation and taphonomy of the skin of Haestasaurus becklesii, and a review of sauropod skin morphology, is published by Pittman et al. (2022).
• A study on the anatomy and phylogenetic affinities of Ligabuesaurus leanzai, based on data from new postcranial elements assigned to the holotype specimen and from a newly referred specimen, is published by Bellardini et al. (2022).
• Description of teeth of a sauropod belonging to the group Somphospondyli from the Turonian Tamagawa Formation (Japan), and a study on the diet and mastication of this sauropod as inferred from tooth wear, is published by Sakaki et al. (2022).
• A study on the phylogenetic relationships of titanosaur sauropods is published by Carballido et al. (2022).
• A study on the morphological variability of hindlimb bones of titanosaur sauropods from the Lo Hueco Konzentrat-Lagerstätte (Villalba de la Sierra Formation, Spain) is published by Páramo et al. (2022).
• Titanosaur tracks preserving claw impressions are reported from the Anacleto Formation (Argentina) by Tomaselli et al. (2022), who devise a new classification for titanosaur tracks and name the new ichnotaxon Teratopodus malarguensis.
• The first titanosaur nesting site from the Late Cretaceous of Brazil is reported from the Maastrichtian Serra da Galga Formation by Fiorelli et al. (2022).
• Pathological multi-shelled egg is described from a titanosaur nest from the Upper Cretaceous Lameta Formation (India) by Dhiman, Verma & Prasad (2022), who interpret this finding as possible evidence that titanosaurs had an oviductal functional morphology similar to birds.
• Review of the fossil record of titanosaur sauropods from the Campanian and Maastrichtian of South America is published by Santucci & Filippi (2022).
• A juvenile specimen of Diamantinasaurus matildae, providing information on the growth pattern of this sauropod, is described from the Upper Cretaceous Winton Formation (Australia) by Rigby et al. (2022).
• A mechanical analysis of Savannasaurus elliottorum is performed by Preuschoft (2022).
• A review of sauropod fossil material from the Kallamedu Formation, including bones of the giant enigmatic titanosaur Bruhathkayosaurus, is published by Pal & Ayyasami (2022).
• A reconstruction of the articular cartilage of the left elbow joint of Dreadnoughtus schrani is presented by Voegele et al. (2022).
• A study on the microstructure of axial bones of Austroposeidon magnificus, Gondwanatitan faustoi and Maxakalisaurus topai, and on its implications for the knowledge of growth phases of these sauropods, is published by Brum et al. (2022).
• Curved, pencil-like sauropod teeth from the Upper Cretaceous Bostobe Formation (Kazakhstan) are referred to a representative of the clade Opisthocoelicaudiidae by Averianov & Lopatin (2022).
• A study proposing a method to determine the gait and limb phase of sauropods based on fossil tracksites is published by Lallensack & Falkingham (2022), who interpret their findings as suggestive of diagonal couplet walks, which would have allowed both sides of the body to be supported by the limbs at all times.
• Revision of the fossil record of sauropodomorph eggs, nests and embryos from South America is published by Fernández, Vila & Moreno-Azanza (2022).
• Keller & Or (2022) hypothesize that sauropods must have compacted the subsoil during their locomotion, presenting a paradox for productivity of the land that supported them.

Theropod research

• Review of the morphology and distribution of non-feather integumentary structures in non-avialan theropods is published by Hendrickx et al. (2022).
• Description of a small high-density assemblage of theropod tracks from the Cretaceous Haman Formation (South Korea), and a study on the distribution of grallatorid tracks in east Asia, is published by Lockley et al. (2022).
• Trackway produced by a large theropod, probably affected by a foot pathology, is described from the upper Barremian locality of Las Hoyas (La Huérguina Formation, Spain) by Herrera-Castillo et al. (2022).
• Revision of the fossil material of theropods from the Middle to Late Jurassic of the Vaches Noires cliffs (Normandy, France) is published by Monvoisin et al. (2022).
• Revision of theropod teeth from the Campanian site of Laño (Spain), evaluating their implications for the knowledge of diversity and evolutionary history of theropods from the Late Cretaceous of Europe, is published by Isasmendi et al. (2022).
• A study aiming to determine the causes of the shortening of the forelimbs of giant theropods, especially tyrannosaurids, is published by Padian (2022).
• Caudal vertebra of a theropod is described from the Aliança Formation (Brazil) by De Oliveira, Oliveira & Fambrini (2022), who consider the studied specimen to be a basal neotheropod, and interpret this finding as likely evidence of the survival of basal neotheropods into the Middle-Late Jurassic in Gondwana.
• The first definitive fossil (a vertebra) of an abelisaurid from the Upper Cretaceous Bahariya Formation (Egypt) is described by Salem et al. (2022).
• A small abelisaurid caudal vertebra is described from the Upper Cretaceous Presidente Prudente Formation (Brazil) by Delcourt & Langer (2022), who interpret this vertebra as belonging to an adult animal, representing one of the smallest known abelisaurids.
• A study on the bone histology of the type specimen of Aucasaurus garridoi is published by Baiano & Cerda (2022).
• An analysis of the possible aquatic habits of members of Spinosauridae, as well as other non-avian dinosaurs, is published by Fabbri et al., who determine that a high bone density would have allowed for underwater foraging in Spinosaurus and Baryonyx, while Suchomimus was likely better suited for terrestrial wading, despite morphological similarities to Baryonyx.
• Isasmendi et al. (2022) reinterpret a fragment of a maxilla from the Lower Cretaceous of La Rioja (Spain), previously assigned to Baryonyx, as likely belonging to an indeterminate baryonychine closer to Baryonyx than to Suchomimus.
• Postcranial material of a giant spinosaurid, which was likely one of the largest European theropods reported to date, is described from the Lower Cretaceous Vectis Formation (United Kingdom) by Barker et al. (2022).
• Redescription of the first theropod tooth discovered in Australia (probably from the Griman Creek Formation) is published by Kotevski and Poropat (2022), who interpret the tooth as belonging to a member of Megaraptoridae.
• Description of five theropod teeth assignable to three different families (Troodontidae, Dromaeosauridae, and Tyrannosauridae) from the Early Campanian Nenjiang Formation (China) is published by Yu et al. (2022)
• Partial tyrannosauroid femur, morphologically similar to the femur of Moros intrepidus but not referable to this taxon, is described from the Albian–Cenomanian Wayan Formation (Idaho, United States) by Krumenacker, Zanno & Sues (2022), who interpret this finding as evidence of the presence of a previously unrecognized tyrannosauroid in the early Late Cretaceous of Laramidia.
• Two juvenile specimens of Gorgosaurus libratus, providing new information on the anatomy and ontogeny of this taxon and tyrannosaurids in general, are described from the Late Cretaceous Dinosaur Park Formation (Alberta, Canada) by Voris et al. (2022).
• Description of the frontal anatomy of Teratophoneus curriei is published by Yun (2022).
• A study on the anatomy of the skull of Qianzhousaurus sinensis is published by Foster et al. (2022).
• Kim et al. (2022) compare a fish centrum found with the holotype of Raptorex kriegsteini with Harenaichthys lui from the Nemegt Formation (Mongolia) and Chinese Xixiaichthys tongxinensis, and interpret their findings as supporting the conclusion that the holotype of R. kriegsteini comes from the Nemegt Formation.
• Description of the neurovascular canals in rostral cranial elements of Tyrannosaurus rex, and a study on the evolution of these canals among Sauropsida and on the possibility of the presence of lips and specialised sensory organs among non-avian theropods, is published by Bouabdellah, Lessner & Benoit (2022).
• An ornithomimosaurian pelvis and sacrum is described from the Upper Cretaceous Erlian Formation (China) by Xi et al. (2022), who interpret this fossil material as likely belonging to a member of Ornithomimosauria distinct from Archaeornithomimus asiaticus, probably representing an early-diverging group within Ornithomimosauria.
• The first diagnostic ornithomimid fossils from the upper Maastrichtian Scollard Formation (Alberta, Canada) are described by Nottrodt (2022), extending the stratigraphic ranges of both Ornithomimus and Struthiomimus in Alberta from the upper Campanian Dinosaur Park Formation through to the Scollard Formation, which constitutes more than 10 million years of time.
• Redescription of Parvicursor remotus is published by Averianov & Lopatin (2022), who reinterpret the holotype of this genus as a juvenile and consider Linhenykus monodactylus and Ceratonykus oculatus to be synonymous with it.
• A study on the jaw adductor musculature and bite force of members of Oviraptorosauria is published by Meade & Ma (2022).
• Review of the knowledge of the reproductive biology of the Late Cretaceous oviraptorosaurs is published by Yang & Sander (2022).
• A subadult oviraptorid specimen interpreted as the first non-hatchling specimen of Yulong mini reported to date is described from the Upper Cretaceous Qiupa Formation (China) by Wei et al. (2022).
• A study on the evolution of the skull morphology of non-avialan paravian theropods is published by Pei & Xu (2022).
• A dromaeosaurid-like sickle claw, similar in some ways to Pyroraptor olympius, is reported from the Grès à Reptiles Formation (France) by Brilhante et al. (2022).
• A study on the phylogenetic relationships of members of Eudromaeosauria is published by Powers et al. (2022), who interpret Acheroraptor temertyorum and Atrociraptor marshalli as members of the Saurornitholestinae.
• New theropod assemblage, including the first records of a large carcharodontosaur allosauroid and of a troodontid maniraptoran in Appalachia reported to date, as well as the earliest occurrence of a tyrannosauroid in Appalachia reported to date, is described from the Cenomanian Lewisville Formation (Woodbine Group; Texas, United States) by Noto et al. (2022).

Birds

New bird taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aegotheles zealandivetus	Sp. nov	Valid	Worthy et al.	Early Miocene	Bannockburn Formation	New Zealand	An owlet-nightjar; a species of Aegotheles.
Allgoviachen	Gen. et sp. nov	In press	Mayr, Lechner & Böhme	Miocene (Tortonian)		Germany	A member of the family Anatidae. The type species is A. tortonica.
Annakacygna	Gen. et 2 sp. nov	Valid	Matsuoka & Hasegawa	Miocene	Haraichi Formation	Japan	A member of the family Anatidae belonging to the tribe Cygnini. The type species is A. hajimei; genus also includes A. yoshiiensis.
Beiguornis	Gen. et sp. nov	Valid	Wang et al.	Early Cretaceous	Longjiang Formation	China	A member of Enantiornithes. The type species is B. khinganensis.
Dryornis hatcheri	Sp. nov	Valid	Degrange	Miocene	Santa Cruz Formation	Argentina	A member of Cathartidae; a species of Dryornis.
Gypaetus georgii	Sp. nov	In press	Sánchez-Marco	Late Miocene		Spain	A vulture; a species of Gypaetus (bearded vulture).
Miosurnia	Gen. et sp. nov		Li, Stidham & Zhou in Li et al.	Late Miocene	Liushu Formation	China	A true owl belonging to the clade Surniini. The type species is M. diurna.
Miotadorna catrionae	Sp. nov	Valid	Tennyson et al.	Miocene (Altonian)	Bannockburn Formation	New Zealand	A member of the family Anatidae belonging to the subfamily Tadorninae.
Musivavis	Gen. et sp. nov	Valid	Wang et al.	Early Cretaceous (Aptian)	Jiufotang Formation	China	A member of Enantiornithes. The type species is M. amabilis.
Neophron lolis	Sp. nov	In press	Sánchez-Marco	Late Miocene		Spain	A vulture; a species of Neophron (Egyptian vulture).
Pliogallus csarnotanus	Sp. nov	Valid	Kessler & Horváth	Pliocene		Hungary	A member of the family Phasianidae.
Zealandornis	Gen. et sp. nov	Valid	Worthy et al.	Early Miocene	Bannockburn Formation	New Zealand	A bird with morphology most similar to that of mousebirds, assigned to the new family Zealandornithidae of uncertain affinities but likely belonging to Telluraves. The type species is Z. relictus.

Avian research

• A study aiming to determine whether the flapping flight of birds evolved through the stage of wing-assisted incline running is published by Kuznetsov & Panyutina (2022).
• A study on the skeletal morphometrics of a sample of specimens of Confuciusornis sanctus is published by Marugán-Lobón & Chiappe (2022), who interpret their findings as indicating that the polyphasic life cycle of C. sanctus was different from the life cycle of modern birds, and possibly indicative of change of food resources foraged by this bird during its ontogeny.
• Wang et al. (2022) reconstruct the pectoral girdles of Sapeornis and Piscivorenantiornis.
• A study on the diet of members of the family Longipterygidae is published by Miller et al. (2022).
• Review of the general anatomy, taxonomy, phylogeny, evolutionary trends and paleoecology of hesperornithiforms is published by Bell & Chiappe (2022).
• Review of the palaeognath fossil record is published by Widrig & Field (2022).
• A study on the stratigraphic provenance of Psammornis eggshells (probably produced by giant ostriches), and on their implications for the knowledge of the evolutionary history of struthionids, is published by Buffetaut (2022).
• An overview and update of the rhea fossil record from South America and Antarctica is published by Picasso, Acosta Hospitaleche & Mosto (2022).
• A study on the taxonomic identity of the extinct giant bird that produced eggs known from eggshell fragments from Pleistocene sites in Australia, some of which bear signs of cooking during a narrow temporal window 50 ± 5 ka B.P., is published by Demarchi et al. (2022), who interpret their findings as supporting the attribution of the studied eggshells to Genyornis.
• A study on the relationships between the shape and size of extant waterfowl tarsometatarsi and their locomotory habits, and on their implications for the knowledge of the locomotory habits of Cayaoa and Paranyroca, is published by De Mendoza & Gómez (2022).
• A study on the morphological variation of the postcranial skeletons of the Malagasy shelduck, and on its implications for the knowledge of the biology of this species, is published by Nomenjanahary et al. (2022).
• A catalogue of fossil and subfossil birds from Cuba is published by Suárez (2022).
• A new partial pedal phalanx of a large, recently extinct barn-owl is described from the Holocene of Guadeloupe by Gala, Laroulandie, and Lenoble (2022).
• An atlas of a great grey owl is described from the Pleistocene of the Devetashka Cave (Bulgaria) by Boev & Mikkola (2022).
• Description of a partial humerus belonging to an auk from the Pliocene Fukagawa Group (Japan) is published by Aotsuka & Endo (2022).
• Fossil material of buttonquails is described from the latest Oligocene and late early to middle Miocene of France by De Pietri et al. (2022), bridging the large temporal gap in the fossil record of this group from the early Oligocene to the late Miocene.
• A review of the evolutionary and biogeographic history of penguins is published by Pelegrín & Acosta Hospitaleche (2022).
• Description of a new partial fossil sternum belonging to a member of Procellariidae from the Middle Pleistocene Ichijiku Formation (Japan) is published by Aotsuka, Isaji, and Endo (2022).
• Revision of the fossil material of birds from Cooper's D locality (South Africa) is published by Pavia et al. (2022).
• A study on the effect of past climate oscillations in the Western Palearctic and Africa on the distribution of the grey partridge, the common quail, the corn crake, the little owl, the snowy owl and the Alpine chough is published by Carrera, Pavia & Varela (2022).
• A study on plant material from rock overhangs from mid-late Holocene sites along the Kawarau-Cromwell-Roxburgh Gorges in Central Otago (New Zealand), much of which was likely transported as roosting material or consumed by moa birds, and on its implications for the knowledge of moa diet and ecology (including the first known evidence of the consumption of kōwhai by moa birds), is published by Pole (2022).
• Evidence from ancient mitochondrial genomes indicative of increase in the population size and genetic diversity of eastern moa after the Last Glacial Maximum, as well as indicative of higher genetic diversity in eastern moa from the southern extent of their range, is presented by Verry, Mitchell & Rawlence (2022), who interpret their findings as indicating that eastern moa expanded from a single glacial refugium following the Last Glacial Maximum.
• A study on the bone histology of extant and extinct large terrestrial birds, aiming to determine whether bone microanatomy can be used to infer the locomotion mode of large terrestrial birds, is published by Canoville, Chinsamy & Angst (2022).

Pterosaurs

New pterosaur taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Cascocauda	Gen. et sp. nov	Valid	Yang et al.	Middle–Late Jurassic (Callovian–Oxfordian)	Tiaojishan Formation	China	An anurognathid. The type species is C. rong.
Dearc	Gen. et sp. nov	Valid	Jagielska et al.	Middle Jurassic (Bathonian)	Lealt Shale Formation	United Kingdom ( Scotland)	A large (2.5 metre wingspan) rhamphorhynchine pterosaur. Genus includes new species D. sgiathanach.
Elektopterus	Gen. et sp. nov	In press		Late Cretaceous(Cenomanian)	Burmese amber	Myanmar	A late surviving Anurognathid. The type species is E. burmae.	File:Anurognathid in amber backlit.jpg
Pachagnathus	Gen. et sp. nov	Valid	Martínez et al.	Late Triassic (Norian)	Quebrada del Barro Formation	Argentina	A raeticodactylid pterosaur. The type species is P. benitoi.
Thanatosdrakon	Gen. et sp. nov	In press	Ortiz David, González Riga & Kellner	Late Cretaceous (Coniacian–Santonian)	Plottier Formation	Argentina	An azhdarchid. The type species is T. amaru.
Yelaphomte	Gen. et sp. nov	Valid	Martínez et al.	Late Triassic (Norian)	Quebrada del Barro Formation	Argentina	A raeticodactylid pterosaur. The type species is Y. praderioi.

Pterosaur research

• Description of the skeletal anatomy of Dimorphodon macronyx is published by Sangster (2022).
• A study reinterpreting the orbital, antorbital and narial fenestrae in the skulls of the anurognathid pterosaurs, based mainly on data from the skulls of specimens of Batrachognathus volans, and aiming to determine the phylogenetic affinities of anurognathids is published by Dalla Vecchia (2022).
• Two specimens of Kunpengopterus sinensis preserved with bromalites are described from the Jurassic Tiaojishan Formation (China) by Jiang et al. (2022), who interpret the bromalites as fossilized gastric pellets, and evaluate their implications for the knowledge of the diet and the digestive system of this pterosaur.
• A new ctenochasmatid assemblage from the Early Cretaceous Quebrada Monardes Formation (Chile) is reported by Alarcón et al. (2022).
• Redescription of the holotype specimen of Moganopterus zhuiana is published by Gao et al. (2022).
• A study on the soft tissues and on the feasibility of the water launch in an aurorazhdarchid specimen from the Late Jurassic Solnhofen Lagoon is published by Pittman et al. (2022).
• New fossil material of Bogolubovia orientalis is described from the Campanian Rybushka Formation (Penza Oblast, Russia) by Averianov & Kurin (2022), who consider Bogolubovia to be a valid taxon that can be distinguished from the pteranodontids Pteranodon and Volgadraco.
• Redescription and a study on the phylogenetic affinities of Ferrodraco lentoni is published by Pentland et al. (2022).
• New skeleton of Sinopterus, providing additional information of the postcranial morphology of this pterosaur, is described by Zhou, Niu & Yu (2022).
• Description of a medium-sized wing skeleton of Sinopterus from the Jiufotang Formation, possibly representing a late juvenile ontogenetic stage, and a study on the histology and life history of this specimen is published by Zhou et al. (2022).
• Cincotta et al. (2022) describe a specimen of Tupandactylus (belonging or related to the species T. imperator) from the Lower Cretaceous Crato Formation (Brazil) associated with skin, monofilaments and branched integumentary structures preserving melanosomes that show tissue-specific geometries (a feature previously known only from theropod dinosaurs), and interpret this finding as indicating that branched integumentary structures in pterosaurs are feathers, as well as providing evidence of deep evolutionary origins of tissue-specific partitioning of melanosome geometry.
• New pterosaur footprint assemblage is described from the Cenomanian Jangdong Formation (South Korea) by Jung et al. (2022), who interpret the studied footprints as possibly produced by small dsungaripteroids, and possibly indicative of gregarious behavior by individuals from different age groups.
• A fragment of the wing metacarpal of a member or a relative of the genus Lonchognathosaurus is described from the Lower Cretaceous Ilek Formation at the Novochernorechensk locality (Krasnoyarsk Territory, Russia) by Averianov et al. (2022), representing the first pterosaur postcranial bone from this formation and the first record of a dsungaripterid from Russia reported to date.
• Cervical vertebra of a large pterosaur is described from the Campanian coastal marine deposits of Izhberda Quarry near the town of Orsk (Orenburg Oblast, Russia) by Averianov, Zverkov & Nikiforov (2022), who interpret this finding as the first record of a giant azhdarchid on the territory of Russia reported to date.

Other archosaurs

Other archosaur taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Maehary	Gen. et sp. nov	Valid	Kellner et al.	Late Triassic (Norian)	Candelária Sequence of the Santa Maria Supersequence	Brazil	Probably an early-diverging member of Pterosauromorpha. The type species is M. bonapartei.

Other archosaur research

• A study on the morphospace occupation of distinct skeletal regions of lagerpetids, aiming to determine which portions of the lagerpetid skeleton are more similar to the anatomy of pterosaurs, is published by Müller (2022).
• Faxinalipterus minimus, originally classified as an early pterosaur, is reinterpreted as a lagerpetid by Kellner et al. (2022).
• A femur of an indeterminate dinosauromorph is described from the Middle Triassic Dinodontosaurus Assemblage Zone (Pinheiros-Chiniquá Sequence, Brazil) by Müller & Garcia (2022), potentially representing the oldest dinosauromorph from South America reported to date.

General research

• Gatesy et al. (2022) propose a standard methodological approach for measuring the relative position and orientation of the major segments of the pelvis and hindlimb of extant and fossil archosaurs in three dimensions.
• A study on the locomotion of extant and extinct archosaurs, reevaluating postulated superiority of early dinosaurs in locomotor function and performance compared with other archosaurs, is published by Cuff et al. (2022).
• Evidence from molecular analyses of modern and fossil skeletal samples, interpreted as indicating that metabolic rates consistent with endothermy are ancestral to ornithodirans, is presented by Wiemann et al. (2022).
• A study on the diversification and body size evolution of terrestrial pan-avian archosaurs along the Triassic and Early Jurassic is published by Langer & Godoy (2022)
• A study on the potential soaring performances of extinct giant birds and pterosaurs is published by Goto et al. (2022).
• A study on the diversity of avian and non-avian theropod tracks from the Aptian Ninesting Creek site (Gething Formation, British Columbia, Canada) is published by Lockley et al. (2022), who report evidence indicating that the diversity of smaller avian and non-avian theropod tracks outnumbers large theropod morphotypes by about 7:1.
• Description of archosaur eggshells from the Late Cretaceous El Gallo Formation (Mexico) and a study on the use of the eggshells to infer the environment of the area is published by Cabrera-Hernández et al. (2022).

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