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] being at the crossroads of several major cultural areals, the ], the ], the ] and ], has a long and involved ].
The land currently known as ] has a long and involved ]. This is partly due to its geographical location at the crossroads of several major cultural areas: the ], the ], the ] and ]. In addition Egypt has experienced several invasions during its long history, including by the ]ites, the ]ns, the ], the ]ns, the ], the ], the ], the ] and the ]. The various conquests over time have made the relationship between Modern ] and ]ians a topic of investigation.
Egypt has experienced several mass migrations and invasions during its history, including by the Canaanites, the ]ns, the ], the ]ns, the ], the ], the ], the ] nd the ]. The various conquests have made the relationship between Modern Egyptians and Ancient Egyptians unclear


==Prehistory== ==Prehistory==
During the Paleolithic the Nile Valley was inhabited by various hunter gatherer populations. About 10,000 years ago the Sahara Desert had a wet phase. People from the surrounding areas moved into the Sahara, and evidence suggests that the populations of the Nile Valley reduced in size. Rock paintings from Algeria at ] reveal that some of the Saharan population were black Africans. It is unclear when Caucasoid populations, such as the ancestors of Berbers, first arrived in North Africa from Eurasia. There are two theories, one is that the ancestors of the ] have been present in North Africa since ] times, the other theory is that the Caucasoid populations only reached North Africa during the ], when they brought domesticated cereals and animals from the Near East. About 5,000 years ago the wet phase of the Sahara came to end. Saharan population retreated to the south towards the Sahel, and East towards the Nile Valley. It is these populations that played a major role in the formation of the Egyptian state as they brought their food crops, sheep, goats and cattle to the Nile Valley.<ref name="keita_natgeo"/><ref>http://www.comp-archaeology.org/WendorfSAA98.html</ref> During the Paleolithic the Nile Valley was inhabited by various hunter gatherer populations. About 10,000 years ago the Sahara Desert had a wet phase. People from the surrounding areas moved from the Sahara, and evidence suggests that the populations of the Nile Valley reduced in size.<ref name="keita_natgeo"></ref> About 5,000 years ago the wet phase of the Sahara came to end. Saharan population retreated to the south towards the ], and East towards the Nile Valley. It was these populations, in addition to ] from the ], that played a major role in the formation of the Egyptian state as they brought their food crops, sheep, goats and cattle to the Nile Valley.<ref name="keita_natgeo"/><ref name="comp-archaeology.org">http://www.comp-archaeology.org/WendorfSAA98.html</ref>


===Predynastic Egypt=== ===Predynastic Egypt===
{{Main|Predynastic Egypt}}
The predynastic period dates to the end of the fourth millenium BC. From about 5000 to 4200BC the ] Culture flourished in ]. This culture has links to Palestine.<ref>Josef Eiwanger: ''Merimde Beni-salame'', In: ''Encyclopedia of the Archaeology of Ancient Egypt''. Compiled and edited by Kathryn A. Bard. London/New York 1999, p. 501-505</ref> The pottery of the Buto Maadi Culture, best known from the site at ] near Cairo, also shows strong connections to South Palestine.<ref>Jürgen Seeher. ''Ma'adi and Wadi Digla''. in: ''Encyclopedia of the Archaeology of Ancient Egypt''. Compiled and edited by Kathryn A. Bard. London/New York 1999, 455-458</ref> The Predynastic period dates to the end of the fourth millennium BC. From about 4800 to 4300BC the ] flourished in ].<ref name="Bogucki">{{cite book |last=Bogucki |first=Peter I. |title=The origins of human society |year=1999 |publisher=Wiley-Blackwell |isbn=1-57718-112-3 |pages=355 }}</ref> This culture, among others, has links to the ].<ref>Josef Eiwanger: ''Merimde Beni-salame'', In: ''Encyclopedia of the Archaeology of Ancient Egypt''. Compiled and edited by Kathryn A. Bard. London/New York 1999, p. 501-505</ref> The pottery of the Buto Maadi culture, best known from the site at ] near Cairo, also shows connections to the southern ].<ref>Jürgen Seeher. ''Ma'adi and Wadi Digla''. in: ''Encyclopedia of the Archaeology of Ancient Egypt''. Compiled and edited by Kathryn A. Bard. London/New York 1999, 455-458</ref>


In Upper Egypt the predynastic Badarian culture was followed by the Naqada culture. The origins of these people is still not fully understood, but their crops came originally from the Near East. There was also significant contact between the peoples of the Naqada culture and the Nubian A-Group people. The origins of the Egyptian dynastic state can be traced to the Naqada culture. In ] the predynastic ] was followed by the ]. The origins of these people is still not fully understood although they seem to be more closely related to the Nubians and North East Africans than with ].<ref name="zakrzewski2007">{{cite journal|first=Sonia |last=Zakrzewski|url=|title= Population continuity or population change: Formation of the ancient Egyptian state|doi=10.1002/ajpa.20569|year=2007|journal=American Journal of Physical Anthropology|volume=132|pages=501–9|pmid=17295300|issue=4}}</ref><ref> by Maria Gatto, archaeology.org</ref>

Some have suggested that the Nubian A-Group conquered the Naqada and then went on to conquer Lower Egypt to begin the dynastic era. However this view is disputed by many scholars. Some studies have however described human remains from both the Naqada and Badarian cultures as clustering with Nubians or Negroids than with Northern Egyptian remains.<ref name="zakrzewski2007">{{cite journal|first=Sonia |last=Zakrzewski|url=http://wysinger.homestead.com/zakrzewski_2007.pdf|title= Population continuity or population change: Formation of the ancient Egyptian state|doi=10.1002/ajpa.20569|year=2007}}</ref>


===Biogeographic origin based on cultural data=== ===Biogeographic origin based on cultural data===
Located in the extreme north-east corner of Africa, ancient Egyptian society was at a crossroads between the African and Near Eastern regions. Early proponents of the ] based their hypothesis on the increased novelty and seemingly rapid change in pre-dynastic pottery and noted trade contacts between ancient Egypt and the Middle East.<ref>Hoffman. "Egypt before the pharaohs: the prehistoric foundations of Egyptian civilization‎", pp267</ref> This is no longer the dominant view in Egyptology, however the evidence on which it was based still suggests influence from these regions.<ref> Redford, Egypt, Israel, p. 17. </ref> Fekri Hassan and Edwin et al point to mutual influence from both inner Africa as well as the Levant.<ref>Edwin C. M et. al, "Egypt and the Levant", pp514</ref> However according to one author this influence seems to have had minimal impact on the indigenous populations already present.<ref>Toby A.H. Wilkinson. " Early Dynastic Egypt: Strategies, Society and Security‎", pp.15</ref>


Located in the extreme north-east corner of Africa, Ancient Egyptian society was at a crossroads between the African and Near Eastern regions. Early proponents of the ] based their hypothesis on the increased novelty and seemingly rapid change in Predynastic pottery and noted trade contacts between ancient Egypt and the Middle East.<ref>Hoffman. "Egypt before the pharaohs: the prehistoric foundations of Egyptian civilization", pp267</ref> This is no longer the dominant view in Egyptology, however the evidence on which it was based still suggests influence from these regions.<ref>Redford, Egypt, Israel, p. 17.</ref> Fekri Hassan and Edwin et al. point to mutual influence from both inner Africa as well as the Levant.<ref>Edwin C. M et. al, "Egypt and the Levant", pp514</ref> However according to one author this influence seems to have had minimal impact on the indigenous populations already present.<ref>Toby A.H. Wilkinson. " Early Dynastic Egypt: Strategies, Society and Security", pp.15</ref>
One author has stated that the ] phase of predynastic Egyptians in ] shared an almost identical culture with A-group peoples of the Lower Sudan.<ref> - by Maria Gatto, archaeology.org</ref> Based in part on the similarities at the royal tombs at Qustul, some scholars have even proposed an Egyptian origin in Nubia among the A-group.<ref> - Encyclopedia of Precolonial Africa, by Joseph O. Vogel, AltaMira Press, (1997), pp. 465-472</ref><ref>Journal of Near Eastern Studies, Vol. 46, No. 1 (Jan., 1987), pp. 15-26</ref> In 1996 Lovell and Prowse reported the presence of individual rulers buried at Naqada in what they interpreted to be elite, high status tombs, showing them to be more closely related morphologically to populations in Northern Nubia than those in Southern Egypt.<ref>Tracy L. Prowse, Nancy C. Lovell. , American Journal of Physical Anthropology, Vol. 101, Issue 2, October 1996, Pages: 237-246</ref> Most scholars however, have rejected this hypothesis and cite the presence of royal tombs that are contemporaneous with that of Qustul and just as elaborate, together with problems with the dating techniques.<ref>Wegner, J. W. 1996. Interaction between the Nubian A-Group and Predynastic Egypt: The Significance of the Qustul Incense Burner. In T. Celenko, Ed., Egypt in Africa: 98-100. Indianapolis: Indianapolis Museum of Art/Indiana University Press.</ref>


One author has stated that the ] phase of Predynastic Egyptians in ] shared an almost identical culture with the ] peoples of the Lower Sudan.<ref> - by Maria Gatto, archaeology.org</ref> Based in part on the similarities at the royal tombs at Qustul, some scholars{{who|date=January 2013}} have even proposed an Egyptian origin in Nubia among the A-group.<ref>"Egypt and Sub-Saharan Africa: Their Interaction" - Encyclopedia of Precolonial Africa, by Joseph O. Vogel, AltaMira Press, (1997), pp. 465-472</ref><ref>Journal of Near Eastern Studies, Vol. 46, No. 1 (Jan., 1987), pp. 15-26</ref> In 1996 Lovell and Prowse reported the presence of individual rulers buried at Naqada in what they interpreted to be elite, high status tombs, showing them to be more closely related morphologically to populations in Northern Nubia than those in Southern Egypt.<ref>Tracy L. Prowse, Nancy C. Lovell. , ], Vol. 101, Issue 2, October 1996, Pages: 237-246</ref> Most scholars however, have rejected this hypothesis and cite the presence of royal tombs that are contemporaneous with that of ] and just as elaborate, together with problems with the ].<ref>Wegner, J. W. 1996. Interaction between the Nubian A-Group and Predynastic Egypt: The Significance of the Qustul Incense Burner. In T. Celenko, Ed., Egypt in Africa: 98-100. Indianapolis: Indianapolis Museum of Art/Indiana University Press.</ref>
The language of the Nubian people is one of the ], whereas the language of the Egyptian people was one of the Afro-Asiatic languages.


Toby Wilkonson, in his book "Genesis of the Pharaohs", proposes an origin for the Egyptians somewhere in the ].<ref> - Review by Timothy Kendall, American Archaeologist</ref> He presents evidence that much of predynastic Egypt duplicated the traditional African cattle-culture typical of Southern Sudanese and East African pastoralists of today. Kendall agrees with Wilkinson's interpretation that ancient rock art in the region may depict the first examples of the royal crowns, while also pointing to Qustul in Nubia as a likely candidate for the origins of the ], being that the earliest known example of it was discovered in this area. Toby Wilkinson, in his book ''Genesis of the Pharaohs'', proposes an origin for the Egyptians somewhere in the ].<ref> Genesis of the Pharaohs: Genesis of the ‘Ka’ and Crowns?] - Review by Timothy Kendall, American Archaeologist</ref> He presents evidence that much of predynastic Egypt duplicated the traditional African cattle-culture typical of Southern Sudanese and East African pastoralists of today. Kendall agrees with Wilkinson's interpretation that ancient rock art in the region may depict the first examples of the royal crowns, while also pointing to Qustul in Nubia as a likely candidate for the origins of the ], being that the earliest known example of it was discovered in this area.


Excavations from ], located about 100&nbsp;km west of ], suggest that the Neolithic inhabitants of the region were migrants from ]. However there is also evidence that sheep and goats were introduced into Nabta from Southwest Asia about 8000 years ago.<ref> http://www.comp-archaeology.org/WendorfSAA98.html</ref> There is some speculation that this culture is likely to be the predecessor of the Egyptians, based on cultural similarities and social complexity which is thought to be reflective of Egypt's ].<ref></ref><ref>{{cite book |title=Holocene Settlement of the Egyptian Sahara | first=Fred |last=Wendorf| isbn=0306466120|publisher=Springer|date=2001|pages=525|url=http://books.google.com/books?id=qUk0GyDJRCoC&pg=PA525&dq=nabta+playa+sub-saharan&sig=I0g-2CzdwuyL0YG5NJCT0drbHOM }}</ref> There is also evidence that sheep and goats were introduced into ] from ] about 8,000 years ago.<ref name="comp-archaeology.org"/> There is some speculation that this culture is likely to be the predecessor of the Egyptians, based on cultural similarities and social complexity which is thought to be reflective of Egypt's ].<ref></ref><ref>{{cite book |title=Holocene Settlement of the Egyptian Sahara | first=Fred |last=Wendorf| isbn=0-306-46612-0|publisher=Springer|year=2001|pages=525|url=http://books.google.com/?id=qUk0GyDJRCoC&pg=PA525&dq=nabta+playa+sub-saharan }}</ref>


==DNA studies== ===DNA records===
{{see|Archaeogenetics of the Near East}} {{main|DNA history of Egypt}}
Attempts to extract ancient DNA or ] from Ancient Egyptian remains have yielded little or no success. Climatic conditions and the mummification process could hasten the deterioration of DNA. Contamination from handling and intrusion from microbes have also created obstacles to recovery of Ancient DNA.<ref></ref> Consequently most DNA studies have been carried out on modern Egyptian populations with the intent of learning about the influences of historical migrations on the population of Egypt.<ref name="S.O.Y. Keita & A. J. Boyce">{{cite journal|title=Genetics, Egypt, and History: Interpreting Geographical Patterns of Y Chromosome Variation|year=2005|last= S.O.Y. Keita & A. J. Boyce |doi=10.1353/hia.2005.0013|url= http://wysinger.homestead.com/keita.pdf }}</ref><ref name="Shomarka Keita (2005)">{{cite journal|title=Y-Chromosome Variation in Egypt|last=Shomarka Keita (2005)|10.1007/s10437-005-4189-4|url=http://wysinger.homestead.com/African_Archaeological_Revie__June_2005_.pdf}}</ref><ref name="Keita">{{cite journal|title=History in the Interpretation of the Pattern of p49a,f TaqI RFLP Y-Chromosome Variation in Egypt|year=2005|last=Keita|doi=10.1002/ajhb.20428|url=http://wysinger.homestead.com/keita6.pdf}}</ref><ref> </ref> However, there was one notable study of ancient mummies of the 12th Dynasty, performed by Paabo and Di Rienzo, which identified multiple lines of descent, including some from sub-Saharan Africa.<ref>Paabo, S., and A. Di Rienzo, A molecular approach to the study of Egyptian history. In Biological Anthropology and the Study of Ancient Egypt. V. Davies and R. Walker, eds. pp. 86-90. London: British Museum Press. 1993</ref> The other lineages were not identified but Keita (1996) speculates that they may also have been Negro in origin.<ref>S.O.Y. Keita & A. J. Boyce. Egypt in Africa, (1996), pp. 25-27</ref> Contamination from handling and intrusion from microbes create obstacles to the recovery of ].<ref name="books.google.com"></ref> Consequently most DNA studies have been carried out on modern Egyptian populations with the intent of learning about the influences of historical migrations on the population of Egypt.<ref name="S.O.Y. Keita & A. J. Boyce">{{cite journal|title=Genetics, Egypt, and History: Interpreting Geographical Patterns of Y Chromosome Variation|year=2005|last= ] & A. J. Boyce |doi=10.1353/hia.2005.0013|url= |first1=S. O. Y.|last2=Boyce|first2=A. J. (Anthony J.)|journal=History in Africa|volume=32|issue=1|pages=221|date=June 2009}}</ref><ref name="Shomarka Keita (2005)">{{cite journal|title=Y-Chromosome Variation in Egypt|last=Shomarka Keita (2005)|url=|doi=10.1007/s10437-005-4189-4|year=2005|first1=S. O. Y.|journal=African Archaeological Review|volume=22|issue=2|pages=61}}</ref><ref name="Keita">{{cite journal|title=History in the Interpretation of the Pattern of p49a,f TaqI RFLP Y-Chromosome Variation in Egypt|year=2005|last=Keita|doi=10.1002/ajhb.20428|url=|first1=S.O.Y.|journal=American Journal of Human Biology|volume=17|pages=559–67|pmid=16136533|issue=5}}</ref><ref></ref>


In general, various DNA studies have found that the gene frequencies of modern ] populations are intermediate between those of the ], the ], southern ] and ],<ref>, The intermediacy of North Africa and to a lesser extent ] is apparent</ref> though NRY frequency distributions of the modern Egyptian population appear to be much more similar to those of the Middle East than to any sub-Saharan African population, suggesting a much larger ] genetic component.<ref name="luis">The Levant versus the Horn of Africa: Evidence for Bidirectional Corridors of Human Migrations – Luis; Rowold; Regueiro; Caeiro; Cinnioğlu; Roseman; Underhill; Cavalli-Sforza; and Herrera. – see http://www.pubmedcentral.nih.gov/articlerender.fcgi?artid=1182266</ref><ref name="Cavalli-Sforza">Cavalli-Sforza, L.L., P. Menozzi, and A. Piazza. 1994. The History and Geography of Human Genes. Princeton: Princeton University Press.</ref><ref name="Cavalli-Sforza" /><ref name="Bosch1997">{{cite journal | last1 = Bosch | first1 = E. ''et al.'' | year = 1997 | title = Population history of north Africa: evidence from classical genetic markers | url = | journal = Human Biology | volume = 69 | issue = 3| pages = 295–311 }}</ref><ref>{{cite journal | author=Arredi B, Poloni E, Paracchini S, Zerjal T, Fathallah D, Makrelouf M, Pascali V, Novelletto A, Tyler-Smith C | title=A predominantly Nilo Saharan origin for Y-chromosomal DNA variation in North Africa. | journal=Am J Hum Genet | volume=75 | issue=2 | pages=338–45 | year=2004 | pmid=15202071 | doi = 10.1086/423147 | pmc=1216069}}</ref><ref>{{cite journal | author=Manni F, Leonardi P, Barakat A, Rouba H, Heyer E, Klintschar M, McElreavey K, Quintana-Murci L | title=Y-chromosome analysis in Egypt suggests a genetic regional continuity in Northeastern Africa. | journal=]| volume=74 | issue=5 | pages=645–58 | year=2002 | pmid=12495079 | doi = 10.1353/hub.2002.0054}}</ref><ref>{{cite book|title= The History and Geography of Human Genes|authorlink=Cavalli-Sforza|last=Cavalli-Sforza|chapter=Synthetic maps of Africa|chapterurl=http://books.google.com/books?id=FrwNcwKaUKoC&printsec=frontcover#PPA189,M1|isbn= 0691087504}}The present population of the Sahara is Sudan in the extreme north, with an increase of Negroid component as one goes south</ref>
====DNA studies on modern Egyptians====
Egypt has experienced several mass migrations and invasions during its history, including by the Canaanites, the Libyans, the Nubians, the Assyrians, the Kushites, the Persians, the Greeks, the Romans and the Muslim Arabs. The various conquests have made the relationship between Modern Egyptians and Ancient Egyptians unclear. While Afrocentrists such as ] argue that the Egyptians were primarily ] before the many conquests of Egypt diluted the Africanity of the Egyptian people,<ref>{{cite book|title=Egypt, Child of Africa|authorlink=Ivan van Sertima|chapter=|chapterurl=http://books.google.com/books?id=Y7KmBTz2vUoC&printsec=frontcover#PPA2-IA1,M1|year=1994|isbn=1560007923}}</ref> other scholars such as Frank Yurco believe that Modern Egyptians are the direct descendants of the Ancient Egyptians.<ref>Frank Yurco, "An Egyptological Review" in Mary R. Lefkowitz and Guy MacLean Rogers, eds. ''Black Athena Revisited''. Chapel Hill: University of North Carolina Press, 1996. p. 62-100</ref>


Blood typing and DNA sampling on ancient Egyptian mummies is scant; however, blood typing of dynastic mummies found ABO frequencies to be most similar to modern Egyptians<ref>Borgognini Tarli S.M., Paoli G. 1982. Survey on paleoserological studies. Homo, 33: 69-89</ref> and some also to Northern ] populations. ABO blood group distribution shows that the Egyptians form a sister group to North African populations, including ], ] and ]ers.<ref>Cavalli-Sforza, L.L., P. Menozzi, and A. Piazza. 1994. ''The History and Geography of Human Genes''. Princeton: Princeton University Press, 169-174</ref>
In general, various DNA studies have found that the gene frequencies of modern North African populations are intermediate between those of Sub-Saharan Africa and Eurasia,<ref> The intermediacy of North Africa and to lesser extent East Africa between Africa and Europe is apparent</ref> though possessing a greater genetic affinity with the populations of Eurasia than they do with Sub-Saharan Africans.<ref name="Cavalli-Sforza">Cavalli-Sforza, L.L., P. Menozzi, and A. Piazza. 1994. The History and Geography of Human Genes. Princeton: Princeton University Press. </ref><ref name="Cavalli-Sforza"/><ref name="Bosch1997">Bosch, E. et al. 1997. Population history of north Africa: evidence from classical genetic markers. Human Biology. 69(3):295-311.</ref><ref>{{cite journal | author=Arredi B, Poloni E, Paracchini S, Zerjal T, Fathallah D, Makrelouf M, Pascali V, Novelletto A, Tyler-Smith C | title=A predominantly neolithic origin for Y-chromosomal DNA variation in North Africa. | journal=Am J Hum Genet | volume=75 | issue=2 | pages=338-45 | year=2004 | id=PMID 15202071 | doi = 10.1086/423147}}</ref><ref>{{cite journal | author=Manni F, Leonardi P, Barakat A, Rouba H, Heyer E, Klintschar M, McElreavey K, Quintana-Murci L | title=Y-chromosome analysis in Egypt suggests a genetic regional continuity in Northeastern Africa. | journal=Hum Biol | volume=74 | issue=5 | pages=645-58 | year=2002 | id=PMID 12495079 | doi = 10.1353/hub.2002.0054}}</ref><ref>{{cite book|title= The History and Geography of Human Genes|authorlink=Cavalli-Sforza|last=Cavalli-Sforza|chapter=Synthetic maps of Africa|chapterurl=http://books.google.com/books?id=FrwNcwKaUKoC&printsec=frontcover#PPA189,M1}}The present population of the Sahara is Caucasoid in the extreme north, with a fairly gradual increase of Negroid component as one goes south</ref>


Earlier disputed DNA studies of King ] (from 1332 B.C.) from 2011 resulted in scientists at a Zurich-based DNA genealogy centre, iGENEA, in a Discovery Channel documentary 2011 claiming ] had ], to which more than 50% of European men, but less than 1% of modern-day Egyptians, belong to.<ref>. ], 1 August 2011. Retrieved on 6 August 2011</ref> Although these findings were widely reported, the Y-chromosome of King Tut had never been published and the DNA profile displayed in Discovery Channel documentary may not actually have belonged to the Pharaoh. According to Carsten Pusch, a geneticist at Germany's University of Tübingen who was part of the team that unraveled Tut's DNA from samples taken from his mummy and mummies of his family members, iGENEA's claims are "simply impossible." <ref>, Igenea,</ref><ref>, LiveScience, 3 August 2011</ref> R1b does show up in parts of North Africa, particularly some regions in Algeria, where tests have found it in 11.8% of subjects.<ref name=Robino>{{Cite journal| title=Analysis of Y-chromosomal SNP haplogroups and STR haplotypes in an Algerian population sample| first9=C| last9=Torre| first8=S| last8=Inturri| first7=A| last7=Piazza| first6=N| last6=Cerutti| first5=S| last5=Benhamamouch| first4=A| last4=Bekada| first3=C| last3=Di Gaetano| journal=Journal International Journal of Legal Medicine| first2=F| volume=122|issue=3| last2=Crobu|year=2008| pmid=17909833|doi=10.1007/s00414-007-0203-5| author=Robino et al.| pages=251–5| postscript=<!-- Bot inserted parameter. Either remove it; or change its value to "." for the cite to end in a ".", as necessary. -->}}</ref> It is also found in Central Africa around Chad and Cameroon,<ref>{{Cite journal|pmid=11910562 |pmc=447595 |doi=10.1086/340257 |year=2002 |last1=Cruciani |first1=F |last2=Santolamazza |last3=Shen |last4=Macaulay |last5=Moral |last6=Olckers |last7=Modiano |last8=Holmes |last9=Destro-Bisol |title=A back migration from Asia to Sub-Saharan Africa is supported by high-resolution analysis of human Y-chromosome haplotypes. |volume=70 |issue=5 |pages=1197–214 |journal=American Journal of Human Genetics |first2=P |first3=P |first4=V |first5=P |first6=A |first7=D |first8=S |first9=G}}, pp. 13–14</ref> but the Chadic-speaking area in Africa is dominated by the branch known as R1b1c (R-V88).<ref name=Cruciani2010>{{Cite journal|author=Cruciani et al.|year=2010|journal=European Journal of Human Genetics | doi=10.1038/ejhg.2009.231|title=Human Y chromosome haplogroup R-V88: a paternal genetic record of early mid Holocene trans-Saharan connections and the spread of Chadic languages|pmc=2987365|url=http://www.nature.com/ejhg/journal/vaop/ncurrent/abs/ejhg2009231a.html|pmid=20051990|last2=Trombetta|first2=B|last3=Sellitto|first3=D|last4=Massaia|first4=A|last5=Destro-Bisol|first5=G|last6=Watson|first6=E|last7=Beraud Colomb|first7=E|last8=Dugoujon|first8=JM|last9=Moral|first9=P|volume=18|issue=7|pages=800–7|postscript=<!--None-->}}</ref>
Luis, Rowold et al found that the diverse NRY haplotypes observed in a population of mixed Arabs and Berbers found that the majority of haplogroups, about 59% were of Eurasian origin. They found that markers signaling the Neolithic expansion from the Middle East constitute the predominant component. The remaining 39.5% were clades that belonged to ], the predominant Haplogroup in Sub-Saharan Africa. E3b was the predominant African clade of Haplogroup E found in the Egyptian population. E3b is the haplogroup characteristic of Afro-Asiatic speakers and evidence suggests that E3b originated in East Africa. About 9% of were clades associated with Sub-Saharan Africans who are not Afro-Asiatic speakers.<ref name="luis"> The Levant versus the Horn of Africa: Evidence for Bidirectional Corridors of Human Migrations – Luis; Rowold; Regueiro; Caeiro; Cinnioğlu; Roseman; Underhill; Cavalli-Sforza; and Herrera. – see http://www.pubmedcentral.nih.gov/articlerender.fcgi?artid=1182266</ref>


However "half of the researchers" in the DNA field dispute these conclusions, and claim instead that DNA sequencing from ancient material is unreliable and prone to contamination.<ref>http://www.nature.com/news/2011/110427/full/472404a.html</ref>
A study by Krings et al. from 1999 on ] ] along the Nile Valley found that a Eurasian cline runs from Northern Egypt to Southern Sudan, and a Sub-Saharan cline extends from Southern Sudan to Northern Egypt. <ref name="krings">{{cite journal|year=|last=Krings|title=mtDNA Analysis of Nile River Valley Populations: Genetic Corridor or a Barrier to Migration?|url=http://genapps.uchicago.edu/labweb/pubs/krings.pdf|pmid=PMC1377841}}</ref> Another study based on ] lineages links modern Egyptians with people from modern ]/] such as the ]-speaking ].<ref>{{cite journal | author=Kivisild T, Reidla M, Metspalu E, Rosa A, Brehm A, Pennarun E, Parik J, Geberhiwot T, Usanga E, Villems R | title=Ethiopian mitochondrial DNA heritage: tracking gene flow across and around the gate of tears. | journal=Am J Hum Genet | volume=75 | issue=5 | pages=752-70 | year=2004 | id=PMID 15457403 | doi = 10.1086/425161}}</ref> Similarly, an mtDNA study of modern Egyptians from the ] region near Thebes in Southern Egypt revealed that Eurasian haplogroups represented 61% of the population, with the remainder 39% being of Sub-Saharan origin. The oral tradition of the Gurna people indicates that they descend from the ancient Egyptians <ref name="stevanovitch">{{cite journal|title=Mitochondrial DNA Sequence Diversity in a Sedentary Population from Egypt |year=2004 |doi=10.1046/j.1529-8817.2003.00057.x|url=http://www3.interscience.wiley.com/cgi-bin/fulltext/118745570/HTMLSTART|last=Stevanovitch}}</ref>

A study using the ] of modern Egyptian males found similar results, namely that African haplogroups are predominant in the South but the predominant haplogroups in the North are characteristic of other North African populations.<ref name="lucotte">{{cite journal|title=Brief communication: Y-chromosome haplotypes in Egypt |year=2001|last=Lucotte|doi=10.1002/ajpa.10190|url=http://wysinger.homestead.com/haplotypes_in_egypt.pdf}}</ref>.

A study of ]ic ethnic group in Sudan found relatively high frequencies of Sub-Saharan ]. The Copts are descendants of Egyptians who have recently migrated from Egypt. According to the study, the presence of Sub-Saharan haplogroups is consistent with the historical record in which southern Egypt was colonized by Nilotic populations during the early state formation.<ref name="hassan">{{cite journal|last=Hassan|title=Y-Chromosome Variation Among Sudanese:Restricted Gene Flow, Concordance With Language, Geography, and History|url=http://dirkschweitzer.net/E3b-papers/Hassan-Sudan-2008-AJPA.pdf|year=2008|doi=10.1002/ajpa.20876}}</ref>

The results of these genetic studies is consistent with the historical record, which records significant bidirectional contact between Egypt and Nubia within the last few thousand years.<ref name="krings"/><ref name="lucotte"/>


==Anthropometric indicators== ==Anthropometric indicators==


=== Craniofacial criteria=== === Craniofacial criteria===
] ]] ].]]
In 1912 Franz Boas demonstrated that cranial shape is heavily influenced by environmental factors, and therefore ] cannot be a reliable indicator of inherited influences such as race.<ref>Boas, “Changes in Bodily Form of Descendants of Immigrants” (American Anthropologist 14:530–562, 1912)</ref> This conclusion was supported in 2003 in a paper by Gravlee, Bernard and Leonard.<ref>http://www.anthro.fsu.edu/people/faculty/CG_pubs/gravlee03b.pdf</ref><ref>Clarence C. Gravlee, H. Russell Bernard, and William R. Leonard find in “Heredity, Environment, and Cranial Form: A Re-Analysis of Boas’s Immigrant Data” (American Anthropologist 105:123–136, 2003)</ref> The use of craniofacial criteria as reliable indicators of population grouping or ethnicity are now disputed. In 1912 Franz Boas demonstrated that cranial shape is heavily influenced by environmental factors, and can change within a few generations if conditions change, and therefore ] cannot be a reliable indicator of inherited influences such as ethnicity.<ref>Boas, "Changes in Bodily Form of Descendants of Immigrants" (American Anthropologist 14:530–562, 1912)</ref> This conclusion was supported in 2003 in a paper by Gravlee, Bernard and Leonard.<ref>http://www.anthro.fsu.edu/people/faculty/CG_pubs/gravlee03b.pdf</ref><ref>Clarence C. Gravlee, H. Russell Bernard, and William R. Leonard find in "Heredity, Environment, and Cranial Form: A Re-Analysis of Boas’s Immigrant Data" (American Anthropologist 105:123–136, 2003)</ref> A study by Beals, Smith, and Dodd (1984) found that "race" and cranial variation had low correlations, and that cranial variation was instead strongly correlated with climate variables.<ref>, by Leonard Lieberman</ref> This view is also supported by Kemp.<ref>Ancient Egypt: anatomy of a civilization, by Barry J. Kemp pg 47</ref> Other studies have shown that the typical cranial shapes of some African (Sudanic and Ethiopic), Arab and Berber ethnic groups are largely the same.<ref>"The races of man: an outline of anthropology and ethnography", by Joseph Deniker, pg 432</ref><ref>"Papers on inter-racial problems", by Gustav Spiller, pg 24</ref>


Others suggest that craniometry is a reliable indicator of genetic populations in biological anthropology.<ref>http://physanth.net/2009/05/13/is-craniometry-scientific/</ref><ref>von Cramon-Taubadel N. (2011). The relative efficacy of functional and developmental cranial modules for reconstructing global human population history. American Journal of Physical Anthropology 146:83-93.</ref><ref>von Cramon-Taubadel N. & Lycett SJ. (2008). Human cranial variation fits iterative founder effect model with African origin. American Journal of Physical Anthropology 136: 108-113.</ref><ref>von Cramon-Taubadel N. (2009a). Congruence of individual cranial bone morphology and neutral molecular affinity patterns in modern humans. American Journal of Physical Anthropology 140:205-215.</ref><ref>http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1325007/</ref>
A survey cited by Kemp (2005) of ancient Egyptian crania spanning all time periods found that the Egyptian population as a whole clusters more closely to the ]n and ] groups of the Nile valley than to Asian and Mediterranean groups, but that they also cluster more closely to the Asian and Mediterranean groups than they did to the Negroid and Archaic African groups. Kemp also noted that Egypt conquored and settled Nubia beginning in the 1st Dynasty.<ref name=kemp>{{cite book|last=Kemp|first= Barry|title=Egypt: Anatomy of a civilization|year=2005|chapter=Who were the Ancient Egyptians|chapterurl=http://books.google.com/books?id=l-t5vWHAVN0C&printsec=frontcover#PPA53,M1|pages=52}}</ref>

A craniofacial study by ] et al. (1993) concluded that: "The Predynastic of Upper Egypt and the Late Dynastic of Lower Egypt are more closely related to each other than to any other population. As a whole, they show ties with the European Neolithic, North Africa, modern Europe, and, more remotely, India, but not at all with sub-Saharan Africa, eastern Asia, Oceania, or the New World. Adjacent people in the Nile valley show similarities in trivial traits in an unbroken series from the delta in the north southward through Nubia and all the way to Somalia at the equator. At the same time, the gradient in skin color and body proportions suggests long-term adaptive response to selective forces appropriate to the latitude where they occur. An assessment of “race” is as useless as it is impossible. Neither clines nor clusters alone suffice to deal with the biological nature of a widely distributed population. Both must be used."<ref>Brace et al., 'Clines and clusters versus "race"' (1993)</ref> He also commented, "We conclude that the Egyptians have been in place since back in the ] and have been largely unaffected by either invasions or migrations. As others have noted, Egyptians are Egyptians, and they were so in the past as well." The results of this study, however, have been criticized and contradicted by the works of other anthropologists such as ] and Sonia Zakrzewski.{{citation needed|date=September 2013}}
A survey cited by Kemp (2005) of pooled ancient Egyptian crania spanning all time periods found that the Egyptian population as a whole clusters more closely to ''modern'' Egyptians than to other groups, but apart from modern Egyptians, they cluster closest to Nubian and "Ethiopic" populations than they do to Middle Easterners or Europeans. In Kemp's unpooled dendrogram it details that the Pre-Dynastic Egyptians (El Bardi and Naqada) samples cluster closest to ancient Nubians and modern Ethiopic populations, and conversely that Late Kingdom and modern Egyptians cluster with Middle Eastern and modern European populations. Kemp also noted that Egypt conquered and settled Nubia beginning in the 1st Dynasty.<ref name=kemp>{{cite book|last=Kemp|first= Barry|title=Egypt: Anatomy of a civilization|year=2005|chapter=Who were the Ancient Egyptians|chapterurl=http://books.google.com/books?id=l-t5vWHAVN0C&printsec=frontcover#PPA53,M1|pages=52|isbn=0-415-01281-3}}</ref>


Anthropologist Nancy Lovell states the following: Anthropologist Nancy Lovell states the following:
{{cquote|"There is now a sufficient body of evidence from modern studies of skeletal remains to indicate that the ancient Egyptians, especially southern Egyptians, exhibited physical characteristics that are within the range of variation for ancient and modern indigenous peoples of the Sub Sahara and tropical Africa.. In general, the inhabitants of Upper Egypt and Nubia had the greatest biological affinity to people of the ] and more southerly areas."
<blockquote> "must be placed in the context of hypotheses informed by archaeological, linguistic, geographic and other data. In such contexts, the physical anthropological evidence indicates that early Nile Valley populations can be identified as part of an African lineage, but exhibiting local variation. This variation represents the short and long term effects of evolutionary forces, such as gene flow, genetic drift, and natural selection, influenced by culture and geography." <ref>Nancy C. Lovell, " Egyptians, physical anthropology of," in Encyclopedia of the Archaeology of Ancient Egypt, ed. Kathryn A. Bard and Steven Blake Shubert, ( London and New York: Routledge, 1999). pp 328-332)</ref></blockquote>


"must be placed in the context of hypotheses informed by archaeological, linguistic, geographic and other data. In such contexts, the physical anthropological evidence indicates that early Nile Valley populations can be identified as part of an African lineage, but exhibiting local variation. This variation represents the short and long term effects of evolutionary forces, such as gene flow, genetic drift, and natural selection, influenced by culture and geography."<ref>Nancy C. Lovell, " Egyptians, physical anthropology of," in Encyclopedia of the Archaeology of Ancient Egypt, ed. Kathryn A. Bard and Steven Blake Shubert, ( London and New York: Routledge, 1999). pp 328-332</ref>}}
This view was also shared by the late Egyptologist, Frank Yurco.<ref>Frank Yurco, "An Egyptological Review" in Mary R. Lefkowitz and Guy MacLean Rogers, eds. ''Black Athena Revisited''. Chapel Hill: University of North Carolina Press, 1996. p. 62-100</ref>


This view was also shared by the late Egyptologist Frank Yurco.<ref name="Yurco Athena">Frank Yurco, "An Egyptological Review" in Mary R. Lefkowitz and Guy MacLean Rogers, eds. ''Black Athena Revisited''. Chapel Hill: University of North Carolina Press, 1996. pp. 62–100</ref>
A 2005 study by Keita of predynastic ] (Southern Egyptian) crania found that the Badarian samples cluster more closely with the tropical African samples than with European samples, although no Asian samples were included.<ref name="S.O.Y. Keita (2005)">{{cite journal|title=Early Nile Valley Farmers, From El-Badari, Aboriginals or “European” Agro-Nostratic Immigrants? Craniometric Affinities Considered With Other Data|last= S.O.Y. Keita (2005)|doi= 10.1177/0021934704265912|url= http://wysinger.homestead.com/badari.pdf}}</ref>


A 2005 study by Keita of predynastic ] (Southern Egyptian) crania found that the Badarian samples cluster more closely with East African (Ethiopic) samples than they do with Northern European (Berg and Norse) samples, though importantly no Asian and Southern Africa samples were included in the study.<ref name="S.O.Y. Keita (2005)">{{cite journal|title=Early Nile Valley Farmers, From El-Badari, Aboriginals or "European" Agro-Nostratic Immigrants? Craniometric Affinities Considered With Other Data|last= S.O.Y. Keita (2005)|doi= 10.1177/0021934704265912|url= |year=2005|first1=S. O. Y.|journal=Journal of Black Studies|volume=36|issue=2|pages=191}}</ref> Keita has also said that the predynastic crania are different to the lower Egyptian samples, which display a mean part way between modern Sub Saharans Africans and Ethiopians.
Sonia Zakrzewski in 2007 noted that genetic continuity occurs over the Egyptian Predynastic and Early Dynastic periods, but that a relatively high level of genetic differentiation was sustained over this time period. She concluded therefore that the process of state formation itself may have been mainly an indigenous process, but that it may have occurred in association with in-migration, particularly during the Early Dynastic and Old Kingdom periods.<ref>http://wysinger.homestead.com/zakrzewski_2007.pdf</ref>


Sonia Zakrzewski in 2007 noted that population continuity occurs over the Egyptian Predynastic into the Greco-Roman periods, and that a relatively high level of genetic differentiation was sustained over this time period. She concluded therefore that the process of state formation itself may have been mainly an indigenous process, but that it may have occurred in association with in-migration, particularly during the Early Dynastic and Old Kingdom periods.<ref>Zakrzewski, Sonia "Population Continuity or Population Change:Formation of the Ancient Egyptian State" ''AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY 132:501–509 (2007)''</ref>
However a craniofacial study by ] et. al. (1993) concluded that: "The Predynastic of Upper Egypt and the Late Dynastic of Lower Egypt are more closely related to each other than to any other population. As a whole, they show ties with the European Neolithic, North Africa, modern Europe, and, more remotely, India, Somalia, eastern Asia, Oceania, or the New World."<ref>Brace et al., 'Clines and clusters versus "race"' (1993)</ref>


In 2008 Keita found that the early predynastic groups in Southern Egypt were similar craniometrically to Nile valley groups of Ethiopic extraction, and as a whole the dynastic Egyptians (includes both Upper and Lower Egyptians) show much closer affinities more southerly Northeast African populations. He also concluded that more material is needed to make a firm conclusion about the relationship between the early ] Nile valley populations and later ancient Egyptians.<ref>Keita, S.O.Y. "Temporal Variation in Phenetic Affinity of Early Upper Egyptian Male Cranial Series", Human Biology, Volume 80, Number 2 (2008)</ref>
Anthropologist ] and others have pointed out the apparent contradictions of these conclusions, and have also pointed out that such relationships need not necessarily suggest gene-flow.<ref name="S.O.Y. Keita and Rick A. Kittles">{{cite journal|title=The Persistence of Racial Thinking and the Myth of Racial Divergence|year=1997|last= S.O.Y. Keita and Rick A. Kittles |doi= 10.1525/aa.1997.99.3.534 |url= http://wysinger.homestead.com/racial_thinking_-_keita.pdf }}</ref>

In 2008 Keita again found that the early predynastic/dynastic groups in Southern Egypt were similar craniometrically to African Nilotic groups, but concluded that more material is needed to make a firm conclusion about the relationship between the early ] Nile valley populations and later ancient Egyptians.<ref>Keita, S.O.Y. "Temporal Variation in Phenetic Affinity of Early Upper Egyptian Male Cranial Series", Human Biology, Volume 80, Number 2 (2008)</ref>


===Limb ratios=== ===Limb ratios===
Anthropologist ] points out that limb elongation is "clearly related to the dissipation of metabolically generated heat" in areas of higher ambient temperature. He also stated that "skin color intensification and distal limb elongation is apparent wherever people have been long-term residents of the tropics". These features have been observed among Egyptian samples.<ref>Brace CL, Tracer DP, Yaroch LA, Robb J, Brandt K, Nelson AR (1993). ''''. Yrbk Phys Anthropol 36:1–31'.</ref> According to Robins and Shute the average limb elongation ratios among ancient Egyptians is higher than that of modern West Africans who reside much closer to the equator. Robins and Shute therefore term the ancient Egyptians to be "super-negroid" but state that although the body plans of the ancient Egyptians were closer to those of modern negroes than for modern whites, “this does not mean that the ancient Egyptians were negroes".<ref> - Robins, Gay. Human Evolution, Volume 1, Number 4 / August, 1986</ref> Anthropologist S.O.Y. Keita criticized Robins and Shute, stating they do not interpret their results within an adaptive context, and stating that they imply “misleadingly” that early southern Egyptians were not a "part of the Saharo-tropical group, which included Negroes".<ref>S.O.Y. Keita. Studies and Comments of Ancient Egyptian Biological Relationships". History in Africa, 20: 129-154 (1993)</ref> Gallagher et al also points out that "body proportions are under strong climatic selection and evidence remarkable stability within regional lineages".<ref>Gallagher et al. "Population continuity, demic diffusion and Neolithic origins in central-southern Germany: The evidence from body proportions.", Homo. Mar 3 (2009)</ref> Zakrzewski (2003) studied skeletal samples from the Badarian period to the Middle Kingdom. She confirmed the results of Robins and Shute that Ancient Egyptians in general had "tropical body plans” but that their proportions were actually "super-negroid".<ref name="Zakrzewski2003">{{cite journal|title=Variation in Ancient Egyptian Stature and Body Proportions|year=2003|last= Zakrzewski |doi= 10.1002/ajpa.10223 |url= http://wysinger.homestead.com/egyptian_body_proportions.pdf }}</ref> Anthropologist ] points out that limb elongation is "clearly related to the dissipation of metabolically generated heat" in areas of higher ambient temperature. He also stated that "skin color intensification and distal limb elongation is apparent wherever people have been long-term residents of the tropics". He also points out that the term "super negroid" is inappropriate, as it is also applied to non negroid populations. These features have been observed among Egyptian samples.<ref>Brace CL, Tracer DP, Yaroch LA, Robb J, Brandt K, Nelson AR (1993). ''Clines and clusters versus "race:" a test in ancient Egypt and the case of a death on the Nile]''. Yrbk Phys Anthropol 36:1–31'.</ref> According to Robins and Shute the average limb elongation ratios among ancient Egyptians is higher than that of modern West Africans who reside much closer to the equator. Robins and Shute therefore term the ancient Egyptians to be "super-negroid" but state that although the body plans of the ancient Egyptians were closer to those of modern negroes than for modern whites, "this does not mean that the ancient Egyptians were negroes".<ref> - Robins, Gay. Human Evolution, Volume 1, Number 4 / August 1986</ref> Anthropologist S.O.Y. Keita criticized Robins and Shute, stating they do not interpret their results within an adaptive context, and stating that they imply "misleadingly" that early southern Egyptians were not a "part of the Saharo-tropical group, which included Negroes".<ref>S.O.Y. Keita. Studies and Comments of Ancient Egyptian Biological Relationships". History in Africa, 20: 129-154 (1993)</ref> Gallagher et al. also points out that "body proportions are under strong climatic selection and evidence remarkable stability within regional lineages".<ref>Gallagher et al. "Population continuity, demic diffusion and Neolithic origins in central-southern Germany: The evidence from body proportions.", Homo. Mar 3 (2009)</ref> Zakrzewski (2003) studied skeletal samples from the Badarian period to the Middle Kingdom. She confirmed the results of Robins and Shute that Ancient Egyptians in general had "tropical body plans" but that their proportions were actually "super-negroid".<ref name="Zakrzewski2003">{{cite journal|title=Variation in Ancient Egyptian Stature and Body Proportions|year=2003|last= Zakrzewski |doi= 10.1002/ajpa.10223 |url= |first1=Sonia R.|journal=American Journal of Physical Anthropology|volume=121|pages=219–29|pmid=12772210|issue=3 }}</ref>


Trikhanus (1981) found Egyptians to plot closest to tropical Africans and not Mediterranean Europeans residing in a roughly similar climatic area.<ref>S.O.Y. Keita, History in Africa, 20: 129-154 (1993)</ref> A more recent study compared ancient Egyptian osteology to that of African-Americans and White Americans, and found that the stature of the Ancient Egyptians was more similar to the stature of African-Americans, although it was not identical.<ref>Raxter et al. " (2008).</ref> Trikhanus (1981) found Egyptians to plot closest to tropical Africans and not Mediterranean Europeans residing in a roughly similar climatic area.<ref>S.O.Y. Keita, History in Africa, 20: 129-154 (1993)</ref> A more recent study compared ancient Egyptian osteology to that of African-Americans and White Americans, and found that the stature of the Ancient Egyptians was more similar to the stature of African-Americans, although it was not identical:<ref>Raxter et al. (2008).</ref>

{{cquote|Our results confirm that, although ancient Egyptians are closer in body proportion to modern American Blacks than they are to American Whites, proportions in Blacks and Egyptians are not identical. }}


===Dental morphology=== ===Dental morphology===
Modern studies on ancient Egyptian dentition clusters the Ancient Egyptians with Caucasoids (Europeans, Western Asians) who have small teeth, as opposed to Negroids (Western Sub-Saharan Africans) who have megadont/large teeth.<ref>Irish J.D. (1998). "Diachronic and synchronic dental trait affinities of late and post-pleistocene peoples from North Africa". Homo. 49(2) 138-155</ref><ref>Hanihara T and Ishida H. (2005). "Metric dental variation of major human populations". American Journal of Physical Anthropology, Vol. 128. Issue 2. pp. 287-298.</ref>
A 2006 ] study on the dental ] of ancient Egyptians by Prof. Joel Irish shows dental traits characteristic of indigenous North Africans and to a lesser extent Southwest Asian and southern European populations. Among the samples included in the study is skeletal material from the ], (from the Roman period) which clustered very closely with the ] series of the ] period. All the samples, particularly those of the Dynastic period, were significantly divergent from a neolithic West Saharan sample from Lower Nubia. Biological continuity was also found intact from the dynastic to the post-pharaonic periods. According to Irish:
<blockquote> samples exhibit morphologically simple, mass-reduced ]s that are similar to those in populations from greater North Africa (Irish, 1993, 1998a–c, 2000) and, to a lesser extent, western Asia and Europe (Turner, 1985a; Turner and Markowitz, 1990; Roler, 1992; Lipschultz, 1996; Irish, 1998a).<ref>Irish pp. 10-11</ref></blockquote>


A 2006 ] study on the dental ] of ancient Egyptians by Prof. Joel Irish shows dental traits characteristic of current indigenous ] and to a lesser extent ] and southern ]an populations, but not at all to ] populations. Among the samples included in the study is skeletal material from the ], (from the Roman period) which clustered very closely with the ] series of the ] period. All the samples, particularly those of the Dynastic period, were significantly divergent from a neolithic West Saharan sample from Lower Nubia. Biological continuity was also found intact from the dynastic to the post-pharaonic periods. According to Irish:
Anthropologist ] takes issue with the suggestion of Irish et al that Egyptians and Nubians were not primary descendants of the African epipaleolithic and Neolithic populations. Keita also criticizes them for ignoring the possibility that the dentition of the ancient Egyptians could have been caused by "in situ microevolution" driven by dietary change, rather than by racial admixture.<ref name="S.O.Y. Keita"/>
{{cquote| samples exhibit morphologically simple, mass-reduced ]s that are similar to those in populations from greater North Africa (Irish, 1993, 1998a–c, 2000) and, to a lesser extent, western Asia and Europe (Turner, 1985a; Turner and Markowitz, 1990; Roler, 1992; Lipschultz, 1996; Irish, 1998a).<ref>Irish pp. 10-11</ref>}}


Anthropologist ] takes issue with the suggestion of Irish that Egyptians and Nubians were not primary descendants of the African ] and ] populations. Keita also criticizes him for ignoring the possibility that the dentition of the ancient Egyptians could have been caused by "in situ microevolution" driven by dietary change, rather than by racial admixture.<ref name="S.O.Y. Keita">{{cite journal|title= Studies and Comments on Ancient Egyptian Biological Relationships|year=1995|last= S.O.Y. Keita|doi=10.1007/BF02444602|url=|first1= S. O. Y.|journal= International Journal of Anthropology|volume= 10|issue= 2-3|pages= 107 }}</ref> However Keita himself has observed population continuity from the ] to the present in modern Egyptians.
==Racial aspects==
{{main|Ancient Egyptian race controversy]]
In ], the "race of the Ancient Egyptians" is a notable topic of controversy.


==The language element==
For example, Afrocentrists such as ] argue that the Egyptians were primarily ] before the many conquests of Egypt diluted the Africanity of the Egyptian people.<ref>{{cite book|title=Egypt, Child of Africa|authorlink=Ivan van Sertima|chapter=|chapterurl=http://books.google.com/books?id=Y7KmBTz2vUoC&printsec=frontcover#PPA2-IA1,M1|year=1994|isbn=1560007923}}</ref>. Others believe that Modern Egyptians are the direct descendants of the Ancient Egyptians, with the various foreign migrations having had little impact on the Egyptian population.<ref>Frank Yurco, "An Egyptological Review" in Mary R. Lefkowitz and Guy MacLean Rogers, eds. ''Black Athena Revisited''. Chapel Hill: University of North Carolina Press, 1996. p. 62-100</ref>
{{Main|Egyptian language|Afroasiatic languages|Afro-Asiatic Urheimat}}
], the world's oldest surviving surgical document. Written in ] in Ancient Egypt around 1600 B.C.]]


Ancient Egyptian languages are classified into six major chronological divisions; ], ], ], ], ], and ], the latter of which was still used as a working language until the 18th Century AD, and is still used as a liturgical language by Egyptian ] to this day.<ref name="Bard">{{cite book |last=Bard |first=Kathryn A. |coauthors=Steven Blake Shubert |title=Encyclopedia of the archaeology of ancient Egypt |year=1999 |publisher=Routledge |isbn=0-415-18589-0 |pages=274 }}</ref>
==References==

{{reflist}}
===Origins===
The Ancient Egyptian language has been classified as a member of the ] ]. There is no agreement on when and where these languages originated, though the language is generally believed to have originated somewhere in or near the region stretching from the ] in the ] to northern ], and from the Eastern ] in ] to the ], or ], ] and ].<ref name=blench2006>Blench R (2006) Archaeology, Language, and the African Past, Rowman Altamira, ISBN 0-7591-0466-2, ISBN 978-0-7591-0466-2, http://books.google.be/books?id=esFy3Po57A8C</ref><ref name=Ehret2004>Ehret C, Keita SOY, Newman P (2004) The Origins of Afroasiatic a response to Diamond and Bellwood (2003) in the Letters of SCIENCE 306, no. 5702, p. 1680 {{doi|10.1126/science.306.5702.1680c}} </ref><ref name=bernal1987>Bernal M (1987) Black Athena: the Afroasiatic roots of classical civilization, Rutgers University Press, ISBN 0-8135-3655-3, ISBN 978-0-8135-3655-2. http://books.google.be/books?id=yFLm_M_OdK4C</ref><ref name=bender1997>Bender ML (1997), Upside Down Afrasian, Afrikanistische Arbeitspapiere 50, pp. 19-34</ref><ref name=militarev2005>Militarev A (2005) Once more about glottochronology and comparative method: the Omotic-Afrasian case, Аспекты компаративистики - 1 (Aspects of comparative linguistics - 1). FS S. Starostin. Orientalia et Classica II (Moscow), p. 339-408. http://starling.rinet.ru/Texts/fleming.pdf</ref> The language of the neighbouring Nubian people is one of the ], and is not one of the ].<ref>http://www.ethnologue.com/show_family.asp?subid=2752-16</ref><ref>http://www.ethnologue.com/show_family.asp?subid=52-16</ref>


==See also== ==See also==
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*] *]
*] *]
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==References==
{{reflist|2}}

==Further reading==
*{{cite journal|title=A study of Egyptian craniology from prehistoric to Roman times|last=Morant|first=G. M.|jstor=2332021|year=1925|publisher=Biometrika|pages=1–52|journal=Biometrika|volume=17|issue=1/2}}
*{{cite book|first=|last=MacIver|authorlink=David Randall-MacIver|title=The Ancient Races of the Thebaid |chapter=chapter 9|chapterurl=http://books.google.com/books?id=gYoTAAAAYAAJ&printsec=titlepage#PRA1-PA86,M1|year=1905}}
*{{cite journal|doi=10.1017/S0021853700000037|title=Evidence of the Early Penetration of Negroes into Prehistoric Egypt|jstor=180563|year=1971|last=Strouhal|first=Eugen|journal=Journal of African History|pages=1–9|volume=12|issue=1|publisher=Cambridge University Press}}

==External links==
*


{{DEFAULTSORT:Population History Of Egypt}}
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Revision as of 14:06, 5 January 2014

The land currently known as Egypt has a long and involved population history. This is partly due to its geographical location at the crossroads of several major cultural areas: the Mediterranean, the Middle East, the Sahara and Sub-Saharan Africa. In addition Egypt has experienced several invasions during its long history, including by the Canaanites, the Libyans, the Nubians, the Assyrians, the Kushites, the Persians, the Greeks, the Romans and the Arabs. The various conquests over time have made the relationship between Modern Egyptians and Ancient Egyptians a topic of investigation.

Prehistory

During the Paleolithic the Nile Valley was inhabited by various hunter gatherer populations. About 10,000 years ago the Sahara Desert had a wet phase. People from the surrounding areas moved from the Sahara, and evidence suggests that the populations of the Nile Valley reduced in size. About 5,000 years ago the wet phase of the Sahara came to end. Saharan population retreated to the south towards the Sahel, and East towards the Nile Valley. It was these populations, in addition to Neolithic farmers from the Near East, that played a major role in the formation of the Egyptian state as they brought their food crops, sheep, goats and cattle to the Nile Valley.

Predynastic Egypt

Main article: Predynastic Egypt

The Predynastic period dates to the end of the fourth millennium BC. From about 4800 to 4300BC the Merimde culture flourished in Lower Egypt. This culture, among others, has links to the Levant. The pottery of the Buto Maadi culture, best known from the site at Maadi near Cairo, also shows connections to the southern Levant.

In Upper Egypt the predynastic Badarian culture was followed by the Naqada culture. The origins of these people is still not fully understood although they seem to be more closely related to the Nubians and North East Africans than with northern Egyptians.

Biogeographic origin based on cultural data

Located in the extreme north-east corner of Africa, Ancient Egyptian society was at a crossroads between the African and Near Eastern regions. Early proponents of the Dynastic Race Theory based their hypothesis on the increased novelty and seemingly rapid change in Predynastic pottery and noted trade contacts between ancient Egypt and the Middle East. This is no longer the dominant view in Egyptology, however the evidence on which it was based still suggests influence from these regions. Fekri Hassan and Edwin et al. point to mutual influence from both inner Africa as well as the Levant. However according to one author this influence seems to have had minimal impact on the indigenous populations already present.

One author has stated that the Naqada phase of Predynastic Egyptians in Upper Egypt shared an almost identical culture with the Nubian A-Group peoples of the Lower Sudan. Based in part on the similarities at the royal tombs at Qustul, some scholars have even proposed an Egyptian origin in Nubia among the A-group. In 1996 Lovell and Prowse reported the presence of individual rulers buried at Naqada in what they interpreted to be elite, high status tombs, showing them to be more closely related morphologically to populations in Northern Nubia than those in Southern Egypt. Most scholars however, have rejected this hypothesis and cite the presence of royal tombs that are contemporaneous with that of Qustul and just as elaborate, together with problems with the dating techniques.

Toby Wilkinson, in his book Genesis of the Pharaohs, proposes an origin for the Egyptians somewhere in the Eastern Desert. He presents evidence that much of predynastic Egypt duplicated the traditional African cattle-culture typical of Southern Sudanese and East African pastoralists of today. Kendall agrees with Wilkinson's interpretation that ancient rock art in the region may depict the first examples of the royal crowns, while also pointing to Qustul in Nubia as a likely candidate for the origins of the white crown, being that the earliest known example of it was discovered in this area.

There is also evidence that sheep and goats were introduced into Nabta from Southwest Asia about 8,000 years ago. There is some speculation that this culture is likely to be the predecessor of the Egyptians, based on cultural similarities and social complexity which is thought to be reflective of Egypt's Old Kingdom.

DNA records

Main article: DNA history of Egypt

Contamination from handling and intrusion from microbes create obstacles to the recovery of Ancient DNA. Consequently most DNA studies have been carried out on modern Egyptian populations with the intent of learning about the influences of historical migrations on the population of Egypt.

In general, various DNA studies have found that the gene frequencies of modern North African populations are intermediate between those of the Near East, the Horn of Africa, southern Europe and Sub Saharan Africa, though NRY frequency distributions of the modern Egyptian population appear to be much more similar to those of the Middle East than to any sub-Saharan African population, suggesting a much larger Eurasian genetic component.

Blood typing and DNA sampling on ancient Egyptian mummies is scant; however, blood typing of dynastic mummies found ABO frequencies to be most similar to modern Egyptians and some also to Northern Haratin populations. ABO blood group distribution shows that the Egyptians form a sister group to North African populations, including Berbers, Nubians and Canary Islanders.

Earlier disputed DNA studies of King Tutankhamun (from 1332 B.C.) from 2011 resulted in scientists at a Zurich-based DNA genealogy centre, iGENEA, in a Discovery Channel documentary 2011 claiming Tutankhamun had Haplogroup R1b1a2, to which more than 50% of European men, but less than 1% of modern-day Egyptians, belong to. Although these findings were widely reported, the Y-chromosome of King Tut had never been published and the DNA profile displayed in Discovery Channel documentary may not actually have belonged to the Pharaoh. According to Carsten Pusch, a geneticist at Germany's University of Tübingen who was part of the team that unraveled Tut's DNA from samples taken from his mummy and mummies of his family members, iGENEA's claims are "simply impossible." R1b does show up in parts of North Africa, particularly some regions in Algeria, where tests have found it in 11.8% of subjects. It is also found in Central Africa around Chad and Cameroon, but the Chadic-speaking area in Africa is dominated by the branch known as R1b1c (R-V88).

However "half of the researchers" in the DNA field dispute these conclusions, and claim instead that DNA sequencing from ancient material is unreliable and prone to contamination.

Anthropometric indicators

Craniofacial criteria

Ancient Egyptian skull, thought to be that of Akhenaten.

The use of craniofacial criteria as reliable indicators of population grouping or ethnicity are now disputed. In 1912 Franz Boas demonstrated that cranial shape is heavily influenced by environmental factors, and can change within a few generations if conditions change, and therefore cranial measurements cannot be a reliable indicator of inherited influences such as ethnicity. This conclusion was supported in 2003 in a paper by Gravlee, Bernard and Leonard. A study by Beals, Smith, and Dodd (1984) found that "race" and cranial variation had low correlations, and that cranial variation was instead strongly correlated with climate variables. This view is also supported by Kemp. Other studies have shown that the typical cranial shapes of some African (Sudanic and Ethiopic), Arab and Berber ethnic groups are largely the same.

Others suggest that craniometry is a reliable indicator of genetic populations in biological anthropology.

A craniofacial study by C. Loring Brace et al. (1993) concluded that: "The Predynastic of Upper Egypt and the Late Dynastic of Lower Egypt are more closely related to each other than to any other population. As a whole, they show ties with the European Neolithic, North Africa, modern Europe, and, more remotely, India, but not at all with sub-Saharan Africa, eastern Asia, Oceania, or the New World. Adjacent people in the Nile valley show similarities in trivial traits in an unbroken series from the delta in the north southward through Nubia and all the way to Somalia at the equator. At the same time, the gradient in skin color and body proportions suggests long-term adaptive response to selective forces appropriate to the latitude where they occur. An assessment of “race” is as useless as it is impossible. Neither clines nor clusters alone suffice to deal with the biological nature of a widely distributed population. Both must be used." He also commented, "We conclude that the Egyptians have been in place since back in the Pleistocene and have been largely unaffected by either invasions or migrations. As others have noted, Egyptians are Egyptians, and they were so in the past as well." The results of this study, however, have been criticized and contradicted by the works of other anthropologists such as S.O.Y. Keita and Sonia Zakrzewski.

A survey cited by Kemp (2005) of pooled ancient Egyptian crania spanning all time periods found that the Egyptian population as a whole clusters more closely to modern Egyptians than to other groups, but apart from modern Egyptians, they cluster closest to Nubian and "Ethiopic" populations than they do to Middle Easterners or Europeans. In Kemp's unpooled dendrogram it details that the Pre-Dynastic Egyptians (El Bardi and Naqada) samples cluster closest to ancient Nubians and modern Ethiopic populations, and conversely that Late Kingdom and modern Egyptians cluster with Middle Eastern and modern European populations. Kemp also noted that Egypt conquered and settled Nubia beginning in the 1st Dynasty.

Anthropologist Nancy Lovell states the following:

"There is now a sufficient body of evidence from modern studies of skeletal remains to indicate that the ancient Egyptians, especially southern Egyptians, exhibited physical characteristics that are within the range of variation for ancient and modern indigenous peoples of the Sub Sahara and tropical Africa.. In general, the inhabitants of Upper Egypt and Nubia had the greatest biological affinity to people of the Sahara and more southerly areas." "must be placed in the context of hypotheses informed by archaeological, linguistic, geographic and other data. In such contexts, the physical anthropological evidence indicates that early Nile Valley populations can be identified as part of an African lineage, but exhibiting local variation. This variation represents the short and long term effects of evolutionary forces, such as gene flow, genetic drift, and natural selection, influenced by culture and geography."

This view was also shared by the late Egyptologist Frank Yurco.

A 2005 study by Keita of predynastic Badarian (Southern Egyptian) crania found that the Badarian samples cluster more closely with East African (Ethiopic) samples than they do with Northern European (Berg and Norse) samples, though importantly no Asian and Southern Africa samples were included in the study. Keita has also said that the predynastic crania are different to the lower Egyptian samples, which display a mean part way between modern Sub Saharans Africans and Ethiopians.

Sonia Zakrzewski in 2007 noted that population continuity occurs over the Egyptian Predynastic into the Greco-Roman periods, and that a relatively high level of genetic differentiation was sustained over this time period. She concluded therefore that the process of state formation itself may have been mainly an indigenous process, but that it may have occurred in association with in-migration, particularly during the Early Dynastic and Old Kingdom periods.

In 2008 Keita found that the early predynastic groups in Southern Egypt were similar craniometrically to Nile valley groups of Ethiopic extraction, and as a whole the dynastic Egyptians (includes both Upper and Lower Egyptians) show much closer affinities more southerly Northeast African populations. He also concluded that more material is needed to make a firm conclusion about the relationship between the early Holocene Nile valley populations and later ancient Egyptians.

Limb ratios

Anthropologist C. Loring Brace points out that limb elongation is "clearly related to the dissipation of metabolically generated heat" in areas of higher ambient temperature. He also stated that "skin color intensification and distal limb elongation is apparent wherever people have been long-term residents of the tropics". He also points out that the term "super negroid" is inappropriate, as it is also applied to non negroid populations. These features have been observed among Egyptian samples. According to Robins and Shute the average limb elongation ratios among ancient Egyptians is higher than that of modern West Africans who reside much closer to the equator. Robins and Shute therefore term the ancient Egyptians to be "super-negroid" but state that although the body plans of the ancient Egyptians were closer to those of modern negroes than for modern whites, "this does not mean that the ancient Egyptians were negroes". Anthropologist S.O.Y. Keita criticized Robins and Shute, stating they do not interpret their results within an adaptive context, and stating that they imply "misleadingly" that early southern Egyptians were not a "part of the Saharo-tropical group, which included Negroes". Gallagher et al. also points out that "body proportions are under strong climatic selection and evidence remarkable stability within regional lineages". Zakrzewski (2003) studied skeletal samples from the Badarian period to the Middle Kingdom. She confirmed the results of Robins and Shute that Ancient Egyptians in general had "tropical body plans" but that their proportions were actually "super-negroid".

Trikhanus (1981) found Egyptians to plot closest to tropical Africans and not Mediterranean Europeans residing in a roughly similar climatic area. A more recent study compared ancient Egyptian osteology to that of African-Americans and White Americans, and found that the stature of the Ancient Egyptians was more similar to the stature of African-Americans, although it was not identical:

Our results confirm that, although ancient Egyptians are closer in body proportion to modern American Blacks than they are to American Whites, proportions in Blacks and Egyptians are not identical.

Dental morphology

Modern studies on ancient Egyptian dentition clusters the Ancient Egyptians with Caucasoids (Europeans, Western Asians) who have small teeth, as opposed to Negroids (Western Sub-Saharan Africans) who have megadont/large teeth.

A 2006 bioarchaeological study on the dental morphology of ancient Egyptians by Prof. Joel Irish shows dental traits characteristic of current indigenous North Africans and to a lesser extent Middle Eastern and southern European populations, but not at all to Sub-Saharan populations. Among the samples included in the study is skeletal material from the Hawara tombs of Fayum, (from the Roman period) which clustered very closely with the Badarian series of the predynastic period. All the samples, particularly those of the Dynastic period, were significantly divergent from a neolithic West Saharan sample from Lower Nubia. Biological continuity was also found intact from the dynastic to the post-pharaonic periods. According to Irish:

samples exhibit morphologically simple, mass-reduced dentitions that are similar to those in populations from greater North Africa (Irish, 1993, 1998a–c, 2000) and, to a lesser extent, western Asia and Europe (Turner, 1985a; Turner and Markowitz, 1990; Roler, 1992; Lipschultz, 1996; Irish, 1998a).

Anthropologist Shomarka Keita takes issue with the suggestion of Irish that Egyptians and Nubians were not primary descendants of the African epipaleolithic and Neolithic populations. Keita also criticizes him for ignoring the possibility that the dentition of the ancient Egyptians could have been caused by "in situ microevolution" driven by dietary change, rather than by racial admixture. However Keita himself has observed population continuity from the Pleistocene to the present in modern Egyptians.

The language element

Main articles: Egyptian language, Afroasiatic languages, and Afro-Asiatic Urheimat
The Edwin Smith papyrus, the world's oldest surviving surgical document. Written in Hieratic script in Ancient Egypt around 1600 B.C.

Ancient Egyptian languages are classified into six major chronological divisions; Archaic Egyptian, Old Egyptian, Middle Egyptian, Late Egyptian, Demotic Egyptian, and Coptic, the latter of which was still used as a working language until the 18th Century AD, and is still used as a liturgical language by Egyptian Copts to this day.

Origins

The Ancient Egyptian language has been classified as a member of the Afro-Asiatic language family. There is no agreement on when and where these languages originated, though the language is generally believed to have originated somewhere in or near the region stretching from the Levant in the Near East to northern Kenya, and from the Eastern Sahara in North Africa to the Red Sea, or Southern Arabia, Ethiopia and Sudan. The language of the neighbouring Nubian people is one of the Nilo-Saharan languages, and is not one of the Afro-Asiatic languages.

See also

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Further reading

  • Morant, G. M. (1925). "A study of Egyptian craniology from prehistoric to Roman times". Biometrika. 17 (1/2). Biometrika: 1–52. JSTOR 2332021.
  • MacIver (1905). "chapter 9". The Ancient Races of the Thebaid. {{cite book}}: External link in |chapterurl= (help); Unknown parameter |chapterurl= ignored (|chapter-url= suggested) (help)
  • Strouhal, Eugen (1971). "Evidence of the Early Penetration of Negroes into Prehistoric Egypt". Journal of African History. 12 (1). Cambridge University Press: 1–9. doi:10.1017/S0021853700000037. JSTOR 180563.

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