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| {{For|an explanation of very similar terms|Hominidae}} | |||
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| {{Taxobox | |||
| | name = Hominoidea | |||
| | fossil_range = Late ] - Recent | |||
| | image = Weisshandgibbon tierpark berlin.jpg | |||
| | image_width = 250px | |||
| | image_caption = ] (''Hylobates lar'') | |||
| | regnum = ]ia | |||
| | phylum = ] | |||
| | classis = ]ia | |||
| | ordo = ]s | |||
| | subordo = ] | |||
| | infraordo = ] | |||
| | parvordo = ] | |||
| | superfamilia = '''Hominoidea''' | |||
| | superfamilia_authority = ], 1825 | |||
| | subdivision_ranks = ] | |||
| | subdivision = | |||
| †]<br /> | |||
| †]<br /> | |||
| ]<br /> | |||
| †]<br /> | |||
| ] | |||
| }} | |||
| '''Apes''' are Old World ] ]s, more specifically a clade of tailless ] ]s, belonging to the biological superfamily '''Hominoidea'''. The apes are native to ] and ]. Apes are the largest primates and the ], an ape, is the largest living ] animal. Hominoids are traditionally forest dwellers, although ]s may range into savanna, and the extinct ]s are famous for being savanna inhabitants, inferred from their morphology. Humans inhabit almost every terrestrial habitat. | |||
| Hominoidea contains two ] of living (extant) species: | |||
| *] consists of four genera and sixteen species of ], including the ] and the ]. They are commonly referred to as lesser apes. | |||
| *] consists of ]s, ]s, ]s, ]s and ]s.<ref name=MSW3>{{MSW3 Groves|pages=178–184|id=12100751}}</ref><ref name=Goodman2>{{cite journal | journal = Journal of Molecular Evolution | year = 1990 | volume = 30 | pages = 260–266 | title = Primate evolution at the DNA level and a classification of hominoids | author = M. Goodman, D. A. Tagle, D. H. Fitch, W. Bailey, J. Czelusniak, B. F. Koop, P. Benson, J. L. Slightom | doi = 10.1007/BF02099995 | pmid = 2109087 | issue = 3}}</ref> Alternatively, the hominidae family are collectively described as the great apes.<ref name=Dixson1981p13/><ref name=DawkinsUse1/><ref>{{Citation |last=Grehan |first=J.R. |year=2006 |title=Mona Lisa Smile: The morphological enigma of human and great ape evolution |journal=Anatomical Record |volume=289B |pages=139–157 }}</ref><ref name=Benton2005p371/> | |||
| Members of the superfamily are called '''hominoids''' (not to be confused with "]s" or "]s"). | |||
| Some or all hominoids are also called "'''apes'''". However, the term "ape" is used in several different senses. It has been used as a synonym for "monkey" or for any tailless primate with a humanlike appearance.<ref name=EB11Ape/> Thus the ], a kind of monkey, is popularly called the "Barbary ape" to indicate its lack of a tail. Biologists have used the term "ape" to mean a member of the superfamily Hominoidea other than humans,<ref name=Dixson1981p13/> or more recently to mean all members of the superfamily Hominoidea, so that "ape" becomes another word for "hominoid".<ref name=Benton2005p371/><ref name=DawkinsUse2/> See also ]. | |||
| Except for gorillas and humans, hominoids are agile climbers of trees. Their diet is best described as ] or ], consisting of leaves, nuts, seeds and fruits, including grass seeds, and in most cases other animals, either hunted or scavenged (or farmed in the case of humans), along with anything else available and easily digested. | |||
| Most nonhuman hominoids are rare or ]. The chief threat to most of the endangered species is loss of tropical ] habitat, though some populations are further imperiled by hunting for ]. | |||
| you might want to ask a monkey who it is | |||
| As ] knowledge developed, it became clear that taillessness occurred in a number of different and otherwise unrelated ]. The term "ape" was then used in two different senses, as shown in the 1910 ] entry. Either "ape" was still used for a tailless humanlike primate or it became a synonym for "monkey".<ref name=EB11Ape>{{Citation |last=Anon. |year=1911 |contribution=Ape |title=Encyclopædia Britannica |publication-place=New York |publisher=Encyclopædia Britannica |url=http://www.archive.org/stream/encyclopaediabri02chisrich#page/160/mode/2up |accessdate=10 July 2011 |edition=11th |volume=XIX }}</ref> | |||
| Sir ] was one of the ] who developed the idea that there were "trends" in primate evolution and that the living members of the order could be arranged in a series, leading through "monkeys" and "apes" to humans. Within this tradition, "ape" refers to all the members of the superfamily Hominoidea, except humans.<ref name=Dixson1981p13>{{Citation |last=Dixson |first=A.F. |year=1981 |title=The Natural History of the Gorilla |publication-place=London |publisher=Weidenfeld & Nicholson |isbn=978-0-297-77895-0 }}, p. 13</ref> Thus "apes" are a ] group, meaning that although all the species of apes descend from a common ancestor, the group does not include all the descendants of that ancestor, because humans are excluded.<ref>Definitions of ] vary; for the one used here see e.g. {{Citation |last=Stace |first=Clive A. |year=2010a |title=Classification by molecules: What’s in it for field botanists? |journal=Watsonia |volume=28 |pages=103–122 |url=http://www.watsonia.org.uk/Wats28p103.pdf |accessdate=07 February 2010 }}, p. 106</ref> The diagram below shows the currently accepted evolutionary relationships of the Hominoidea,<ref name=Goodman2/> with the apes marked by a bracket. | |||
| {{Barlabel|style=font-size:90%|labelwidth=7|align=centre | |||
| |size=5 | |||
| |at1=3.5|color1=purple|label1=apes | |||
| |cladogram= | |||
| {{Cladex|style=width:auto | |||
| |label1= Hominoidea | |||
| |1={{Cladex | |||
| |1={{Cladex | |||
| |1={{Cladex | |||
| |1={{Cladex | |||
| |1=humans (genus ''Homo'')|barbegin2=purple | |||
| |2=chimpanzees (genus ''Pan'')|bar2=purple | |||
| }} | |||
| |2=gorillas (genus ''Gorilla'')|bar2=purple | |||
| }} | |||
| |2=orangutans (genus ''Pongo'')|bar2=purple | |||
| }} | |||
| |2=gibbons (family Hylobatidae)|barend2=purple | |||
| }} | |||
| }} | |||
| }} | |||
| The "apes" are traditionally divided further into the "lesser apes" and the "great apes":<ref>{{Harvnb|Dixson|1981|p=16}}</ref> | |||
| {{Barlabel|style=font-size:90%|labelwidth=7|align=centre | |||
| |size=5 | |||
| |at1=3|color1=darkblue|label1=great apes | |||
| |at2=5|color2=green|label2=lesser apes | |||
| |cladogram= | |||
| {{Cladex|style=width:auto | |||
| |label1= Hominoidea | |||
| |1={{Cladex | |||
| |1={{Cladex | |||
| |1={{Cladex | |||
| |1={{Cladex | |||
| |1=humans (genus ''Homo'')|barbegin2=darkblue | |||
| |2=chimpanzees (genus ''Pan'')|bar2=darkblue | |||
| }} | |||
| |2=gorillas (genus ''Gorilla'')|bar2=darkblue | |||
| }} | |||
| |2=orangutans (genus ''Pongo'')|barend2=darkblue | |||
| }} | |||
| |2=gibbons (family Hylobatidae)|barbegin2=green|barend2=green | |||
| }} | |||
| }} | |||
| }} | |||
| In summary, there are three common uses of the term "ape": non-biologists may not distinguish between "monkeys" and "apes", or may use "ape" for any tailless monkey or nonhuman hominoid, whereas biologists traditionally used the term "ape" for all non-human hominoids as shown above. | |||
| In recent years biologists have generally preferred to use only ] groups in classifications,{{Citation needed|date=July 2011}} that is only groups which include ''all'' the descendants of a common ancestor.<ref>Definitions of ] vary; for the one used here see e.g. {{Citation |last=Mishler |first=Brent D |year=2009 |editor-last=Ayala |editor-first=F.J. |editor2-last=Arp |editor2-first=R. |contribution=Species are not Uniquely Real Biological Entities |title=Contemporary Debates in Philosophy of Biology |pages=110–122 |isbn=978-1-4443-1492-2 |accessdate=19 April 2011 |doi=10.1002/9781444314922.ch6 |lastauthoramp=yes }}, p. 114</ref> The superfamily Hominoidea is one such group (or "]"). Some then use the term "ape" to mean all the members of the superfamily Hominoidea. For example, in a 2005 book, Benton wrote "The apes, Hominoidea, today include the gibbons and orang-utan ... the gorilla and chimpanzee ... and humans".<ref name=Benton2005p371>{{Citation |last=Benton |first=Michael J. |year=2005 |title=Vertebrate palaeontology |publisher=Wiley-Blackwell |isbn=978-0-632-05637-8 |url=http://books.google.com/books?id=SyJO3vpCk8AC |accessdate=10 July 2011 }}, p. 371</ref> The group traditionally called "apes" by biologists is then called the "nonhuman apes". | |||
| See the section ] below for a discussion of changes in scientific classification and terminology. | |||
| ==Biology== | |||
| {{Unreferenced section|date=July 2011}} | |||
| The "lesser apes" are the gibbon family, Hylobatidae with sixteen medium-sized species. Their major differentiating characteristic is their long arms, which they use to ] through the trees. As an evolutionary adaptation to this ] lifestyle, their wrists are ]s. The largest of the gibbons, the ], weighs up to {{convert|14|kg|lb|0|abbr=on}}. In comparison, the smallest "great ape" is the ] at {{convert|40|to|65|kg|lb|0|abbr=on}}. | |||
| The "great apes" were formerly treated as the family Pongidae. As noted above, this definition makes the Pongidae ], and does not show that orangutans, gorillas, chimpanzees and humans are all more closely related to one another than any of these four groups are to gibbons. Further, current evidence implies that humans share a common extinct ancestor with the chimpanzee line, from which we separated more recently than the gorilla line. | |||
| The superfamily Hominoidea falls within ], which also includes the Old World monkeys of Africa and Eurasia. Within this group, both families (Hylobatidae and Hominidae) can be distinguished from Old World monkeys by the number of ] on their ]s (hominoids have five—the "Y-5" molar pattern, Old World monkeys have only four in a bilophodont pattern). Hominoids have more mobile shoulder joints and arms due to the dorsal position of the ], broad ribcages that are flatter front-to-back, and a shorter, less mobile spine compared to Old World monkeys, with caudal (tail) vertebrae greatly reduced, resulting in complete tail loss in living species. These are all anatomical adaptations to vertical hanging and swinging locomotion (]), as well as better balance in a ] pose. However, there are also primates in other families that lack tails, and at least one (the ]) that has been known to walk significant distances bipedally. The front skull is characterised by its sinuses, fusion of the frontal bone and ]. | |||
| Although the hominoid fossil record is far from complete, and the evidence is often fragmentary, there is enough to give a good outline of the evolutionary history of humans. The time of the split between humans and other living hominoids used to be thought to have occurred 15 to 20 million years ago. Some species occurring within that time period, such as '']'', used to be considered as ]s, and possible ancestors of humans. Later fossil finds indicated that ''Ramapithecus'' was more closely related to the orangutan, and new biochemical evidence indicated that the last common ancestor of humans and other hominins occurred between 5 and 10 million years ago, and probably in the lower end of that range. | |||
| ===Behaviour and cognition=== | |||
| {{Refimprove section|date=July 2011}} | |||
| Although there had been earlier studies, the scientific investigation of behaviour and cognition in nonhuman members of the superfamily Hominoidea expanded enormously during the latter half of the twentieth century. Major studies of behaviour in the field were completed on the three better-known "great apes", for example by ], ] and ] (field work on gibbons and the ] is still relatively underdeveloped). These studies have shown that in their natural environments, the nonhuman hominoids show sharply varying social structure: gibbons are monogamous, territorial pair-bonders, orangutans are solitary, gorillas live in small troops with a single adult male leader, while chimpanzees live in larger troops with bonobos exhibiting promiscuous sexual behaviour. Their diets also vary; gorillas are ]s while the others are all primarily ]s, although the common chimpanzee does some hunting for meat. Foraging behaviour is correspondingly variable. | |||
| All the nonhuman hominoids are generally thought of as highly intelligent, and scientific study has broadly confirmed that they perform outstandingly well on a wide range of cognitive tests – though there is relatively little data on gibbon cognition. The early studies by ] demonstrated exceptional ] abilities in chimpanzees, which Köhler attributed to ]. The ] has been repeatedly demonstrated; more recently, the manufacture of tools has been documented, both in the wild and in laboratory tests. ] is much more easily demonstrated in "great apes" than in other primate species. Almost all the studies in ] have been completed with "great apes", and though there is continuing dispute as to whether they demonstrate real language abilities, there is no doubt that they involve significant feats of learning. Chimpanzees in different parts of Africa have developed tools that are used in food acquisition, demonstrating a form of animal culture.<ref>{{cite book | author = William McGrew|title = Chimpanzee material culture: implications for human evolution | year = 1992}}</ref> | |||
| ===Distinction from monkeys=== | |||
| Apes do not possess a tail, unlike most ]s. Monkeys are more likely to be in trees and use their tails for balance. Apes are considerably larger than monkeys, with the exception of ], which are smaller than some monkeys. Apes are considered to be more intelligent than monkeys, which are considered to have more primitive brains.<ref>The gestural communication of apes and monkeys: Josep Call, Michael Tomasello - 2007</ref> Unlike female monkeys which go through the ], ], including humans, go through a ]. | |||
| ==History of hominoid taxonomy== | |||
| The history of hominoid taxonomy is somewhat confusing and complex. The names of subgroups have changed their meaning over time as new evidence, from fossil discoveries and comparisons of anatomy and DNA sequences, has changed understanding of the relationships between hominoids. There has been a gradual demotion of humans from a special position in the taxonomy to being one branch among many. This history illustrates the growing influence of ] (the science of classifying living things by strict descent) on taxonomy. | |||
| As of 2006, there are eight extant ] of hominoids. They are the four genera in the family Hominidae ('']'' – humans, '']'' – chimpanzees and bonobos, '']'', and '']'' – orangutans), and the four genera in the family Hylobatidae or ]s ('']'', '']'', '']'' and '']'').<ref name=MSW3/> (The genus for the ]s was recently changed from '']'' to ''Hoolock''.<ref name=Mootnick>{{cite journal | last = Mootnick | first = A. | coauthors = Groves, C. P. | year = 2005 | title = A new generic name for the hoolock gibbon (Hylobatidae) | journal = International Journal of Primatology | issue = 26 | pages = 971–976 | doi = 10.1007/s10764-005-5332-4 | volume = 26}}</ref>) | |||
| In 1758, ], relying on second- or third-hand accounts, placed a second species in ''Homo'' along with ''H. sapiens'': ''Homo troglodytes'' ("cave-dwelling man"). It is not clear to which animal this name refers, as Linnaeus had no specimen to refer to, hence no precise description. Linnaeus named the orangutan ''Simia satyrus'' ("satyr monkey"). He placed the three genera ''Homo'', '']'' and ''Lemur'' in the order of Primates. | |||
| The ''troglodytes'' name was used for the chimpanzee by ] in 1775 but moved to the genus ''Simia''. The orangutan was moved to the genus '']'' in 1799 by ]. | |||
| Linnaeus's inclusion of humans in the primates with monkeys and apes was troubling for people who denied a close relationship between humans and the rest of the animal kingdom. Linnaeus's Lutheran archbishop had accused him of "impiety." In a letter to ] dated 25 February 1747, Linnaeus wrote: | |||
| :It is not pleasing to me that I must place humans among the primates, but man is intimately familiar with himself. Let's not quibble over words. It will be the same to me whatever name is applied. But I desperately seek from you and from the whole world a general difference between men and simians from the principles of Natural History. I certainly know of none. If only someone might tell me one! If I called man a simian or vice versa I would bring together all the theologians against me. Perhaps I ought to, in accordance with the law of Natural History.<ref>{{cite web | url = http://linnaeus.c18.net/Letters/display_txt.php?id_letter=L0783 | title = Letter, Carl Linnaeus to Johann Georg Gmelin. Uppsala, Sweden, 25 February 1747 | publisher = Swedish Linnaean Society}}</ref> | |||
| Accordingly, ] in the first edition of his ''Manual of Natural History'' (1779), proposed that the primates be divided into the ] (four-handed, i.e. apes and monkeys) and ] (two-handed, i.e. humans). This distinction was taken up by other naturalists, most notably ]. Some elevated the distinction to the level of ]. | |||
| However, the many affinities between humans and other primates — and especially the "great apes" — made it clear that the distinction made no scientific sense. ] wrote, in '']'': | |||
| :The greater number of naturalists who have taken into consideration the whole structure of man, including his mental faculties, have followed Blumenbach and Cuvier, and have placed man in a separate Order, under the title of the Bimana, and therefore on an equality with the orders of the Quadrumana, Carnivora, etc. Recently many of our best naturalists have recurred to the view first propounded by Linnaeus, so remarkable for his sagacity, and have placed man in the same Order with the Quadrumana, under the title of the Primates. The justice of this conclusion will be admitted: for in the first place, we must bear in mind the comparative insignificance for classification of the great development of the brain in man, and that the strongly marked differences between the skulls of man and the Quadrumana (lately insisted upon by ], ], and others) apparently follow from their differently developed brains. In the second place, we must remember that nearly all the other and more important differences between man and the Quadrumana are manifestly adaptive in their nature, and relate chiefly to the erect position of man; such as the structure of his hand, foot, and pelvis, the curvature of his spine, and the position of his head.<ref>{{cite book | author = ]|title = ] | year = 1871 | isbn = 0-7607-7814-0}}</ref> | |||
| <!-- ...history from 1870 to 1960 goes here... --> | |||
| ===Changes in taxonomy=== | |||
| {| class="wikitable" | |||
| | Until about 1960, the hominoids were usually divided into two families: humans and their extinct relatives in ], all other hominoids in Pongidae.<ref>{{cite journal|author = G. G. Simpson | year = 1945 | title = The principles of classification and a classification of mammals | journal = Bull. Am. Mus. Nat. Hist. | volume = 85 | pages = 1–350 | authorlink = George Gaylord Simpson}}</ref> | |||
| | ] | |||
| |- | |||
| | The 1960s saw the application of techniques from ] to primate taxonomy. Goodman used his 1964 immunological study of serum proteins to propose a division of the hominoids into three families, with the "great apes" in Pongidae and the "lesser apes" (gibbons) in ].<ref name=Goodman>{{cite book | author = M. Goodman | year = 1964 | chapter = Man’s place in the phylogeny of the primates as reflected in serum proteins | editor = S. L. Washburn | title = Classification and human evolution | publisher = Aldine, Chicago|pages = 204–234}}</ref> The trichotomy of hominoid families, however, prompted scientists to ask which family ] first from the common hominoid ancestor. | |||
| | ] | |||
| |- | |||
| | Within the superfamily Hominoidea, gibbons are the ]: this means that the rest of the hominoids are more closely related to each other than any of them are to gibbons. This led to the placing of the "great apes" into the family Hominidae along with humans, by demoting the Pongidae to a subfamily; the Hominidae family now contained the subfamilies ] and ]. Again, the three-way split in Ponginae led scientists to ask which of the three genera is least related to the others. | |||
| | ] | |||
| |- | |||
| | Investigation showed ]s to be the outgroup, but comparing humans to all three other hominid genera showed that African "apes" (chimpanzees and gorillas) and humans are more closely related to each other than any of them are to orangutans. This led to the placing of the African hominoids in the subfamily Homininae, forming another three-way split. This classification was first proposed by M. Goodman in 1974.<ref name=Goodman1>{{cite journal | journal = Annual Review of Anthropology | volume = 3 | issue = 1 | pages = 203–228 | year = 1974 | doi = 10.1146/annurev.an.03.100174.001223 | title = Biochemical Evidence on Hominid Phylogeny | author = M. Goodman}}</ref> | |||
| | ] | |||
| |- | |||
| | To try to resolve the hominine trichotomy, some authors proposed the division of the subfamily Homininae into the tribes Gorillini (African "apes") and Hominini (humans). | |||
| | ] | |||
| |- | |||
| | However, DNA comparisons provide convincing evidence that within the subfamily Homininae, gorillas are the outgroup. This suggests that chimpanzees should be in Hominini along with humans. This classification was first proposed (though one rank lower) by M. Goodman et al. in 1990.<ref name=Goodman2/> See ] for more information on the speciation of humans and "great apes". | |||
| | ] | |||
| |- | |||
| | Later DNA comparisons split the gibbon genus ''Hylobates'' into four genera: '']'', '']'', '']'', and '']''.<ref name=MSW3/><ref name=Mootnick/> | |||
| | ] | |||
| |} | |||
| ==Classification and evolution== | |||
| As discussed above, hominoid taxonomy has undergone several changes. Genetic analysis shows that hominoids diverged from the ]s between 29 million and 34.5 million years ago.<ref>{{cite web |url=http://www.redorbit.com/news/science/117352/apes_monkeys_split_earlier_than_fossils_had_indicated/ |title=Apes, Monkeys Split Earlier Than Fossils Had Indicated/ |author= |date= |work= |publisher= |accessdate= }}</ref> The gibbons split from the rest about 18 mya, and the hominid splits happened 14 mya (''Pongo''), 7 mya (''Gorilla''), and 3-5 mya (''Homo'' & ''Pan'').{{Citation needed|date=July 2008}}<!-- note, this figure of 3-5 mya for the Homo/Pan split differs from the figure in the article ], which says 6.5-7.4 mya or alternatively 5.4-6.3 mya--> | |||
| The families, genera and extant species of hominoids are: | |||
| * '''Superfamily Hominoidea'''<ref name=MSW3/> | |||
| ** Family ]: gibbons ("lesser apes") | |||
| *** Genus '']'' | |||
| **** ] or white-handed gibbon, ''H. lar'' | |||
| **** ], ''H. albibarbis'' | |||
| **** ] or black-handed gibbon, ''H. agilis'' | |||
| **** ] or grey gibbon, ''H. muelleri'' | |||
| **** ], ''H. moloch'' | |||
| **** ] or capped gibbon, ''H. pileatus'' | |||
| **** ] or Mentawai gibbon or bilou, ''H. klossii'' | |||
| *** Genus '']'' | |||
| **** ], ''H. hoolock'' | |||
| **** ], ''H. leuconedys'' | |||
| *** Genus '']'' | |||
| **** ], ''S. syndactylus'' | |||
| *** Genus '']'' | |||
| **** ], ''N. concolor'' | |||
| **** ], ''N. nasutus'' | |||
| **** ], ''N. hainanus'' | |||
| **** ] ''N. siki'' | |||
| **** ], ''N. leucogenys'' | |||
| **** ], ''N. gabriellae'' | |||
| ** Family ]: hominids ("great apes", including humans) | |||
| *** Genus '']'': orangutans | |||
| **** ], ''P. pygmaeus'' | |||
| **** ], ''P. abelii'' | |||
| *** Genus '']'': gorillas | |||
| **** ], ''G. gorilla'' | |||
| **** ], ''G. beringei'' | |||
| *** Genus '']'': humans | |||
| **** ], ''H. sapiens'' | |||
| *** Genus '']'': chimpanzees | |||
| **** ], ''P. troglodytes'' | |||
| **** ], ''P. paniscus'' | |||
| ==Cultural aspects of apes== | |||
| {{Unreferenced section|date=July 2011}} | |||
| Often, "apes" (nonhuman hominoids) are said to be the result of a curse—a Jewish folktale claims that one of the races who built the ] became apes as punishment, while Muslim lore says that the Jews of ] became apes as punishment for fishing on the Sabbath. Some sects of Christianity have folklore that claims that these apes are a symbol of lust and were created by Satan in response to God's creation of humans.{{Citation needed|date=November 2009}} It is uncertain whether any of these references are to any specific apes. All of these concepts date from a period when neither the distinction between apes and monkeys, nor the fact that humans are closely related to chimpanzees, was widely understood, if understood at all. | |||
| ==See also== | |||
| {{Portal|Mammals}} | |||
| *] | |||
| *'']'' from the ] | |||
| *] | |||
| *] | |||
| *] (for notable non-fictional apes) | |||
| *] | |||
| ==Notes and references== | |||
| {{Reflist|refs= | |||
| <ref name=DawkinsUse1>Although Dawkins is clear that he uses "apes" for Hominoidea, he also uses "great apes" in ways which exclude humans. Thus in {{Citation |last=Dawkins |first=R. |year=2005 |title=The Ancestor's Tale |edition=p/b |publication-place=London |publisher=Phoenix (Orion Books) |isbn=978-0-7538-1996-8 }}: "Long before people thought in terms of evolution ... great apes were often confused with humans" (p. 114); "gibbons are faithfully monogamous, unlike the great apes which are our closer relatives" (p. 126).</ref> | |||
| <ref name=DawkinsUse2>{{Harvnb|Dawkins|2005}}; for example "ll apes except humans are hairy" (p. 99), "mong the apes, gibbons are second only to humans" (p. 126).</ref> | |||
| }} | |||
| ==External links== | |||
| {{Sister project links|Ape}} | |||
| {{Wikispecies|Hominoidea|Ape}} | |||
| {{Wikibooks|Dichotomous Key|Hominoidea}} | |||
| *{{cite journal |author=Pilbeam D |authorlink=David Pilbeam |title=Hominoid systematics: The soft evidence |journal=Proc. Natl. Acad. Sci. U.S.A. |volume=97 |issue=20 |pages=10684–6 |year=2000 |month=September |pmid=10995486 |pmc=34045 |doi=10.1073/pnas.210390497 |url=http://www.pnas.org/cgi/pmidlookup?view=long&pmid=10995486}} Agreement between cladograms based on molecular and anatomical data. | |||
| {{apes}} | |||
| {{Use dmy dates|date=July 2011}} | |||
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Revision as of 16:57, 1 June 2012
I Like monkeys