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Bucerotiformes Temporal range: Eocene to present
Eurasian Hoopoe (Upupa epops) (Upupidae)
Western red-billed hornbill (Tockus kempi) (Bucerotidae)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Clade:	Picocoraciae
Order:	Bucerotiformes Fürbringer, 1888
Families
Phoeniculidae Upupidae Bucorvidae Bucerotidae

Bucerotiformes /bjuːˈsɛrətɪfɔːrmiːz/ is an order of birds that contains the hornbills, ground hornbills, hoopoes and wood hoopoes. These birds were previously classified as members of Coraciiformes. The clade is distributed in Africa, Asia, Europe and Melanesia.

Systematics

Recent genetic data show that ground hornbills and Bycanistes form a clade outside the rest of the hornbill lineage. They are thought to represent an early African lineage, while the rest of Bucerotiformes evolved in Asia. The hoopoe subspecies Saint Helena hoopoe and the Madagascar subspecies are sometimes elevated to a full species. The two wood hoopoe genera, Phoeniculus and Rhinopomastus, appear to have diverged about 10 million years ago, so some systematists treat them as separate subfamilies or even separate families.

Extinct Messelirrisoridae and Laurillardiidae families were both considered to be stem groups of a previously categorized Upupiformes order prior to it being subcategorized into Bucerotiformes.

Taxonomy

Order Bucerotiformes

• Family †Messelirrisoridae
  • Genus †Messelirrisor
    • †Messelirrisor halcyrostris
    • †Messelirrisor grandis
    • †Messelirrisor parvus
• Family †Laurillardiidae
  • Genus †Laurillardia
    • †Laurillardia longirostris
    • †Laurillardia munieri
    • †Laurillardia smoleni
• Suborder Upupi
  • Family Phoeniculidae
    • Genus Phoeniculus
      • Green wood hoopoe, Phoeniculus purpureus
      • Violet wood hoopoe, Phoeniculus damarensis
      • Black-billed wood hoopoe, Phoeniculus somaliensis
      • White-headed wood hoopoe, Phoeniculus bollei
      • Forest wood hoopoe, Phoeniculus castaneiceps
    • Genus Rhinopomastus
      • Black scimitarbill, Rhinopomastus aterrimus
      • Common scimitarbill, Rhinopomastus cyanomelas
      • Abyssinian scimitarbill, Rhinopomastus minor
  • Family Upupidae
    • Genus Upupa
      • Eurasian hoopoe, Upupa epops
      • African hoopoe, Upupa africana
      • Madagascar hoopoe, Upupa marginata
• Suborder Buceroti
  • Family Bucorvidae
    • Genus Bucorvus
      • Abyssinian ground hornbill, Bucorvus abyssinicus
      • Southern ground hornbill, Bucorvus leadbeateri
  • Family Bucerotidae
    • Genus Bycanistes
      • Trumpeter hornbill, Bycanistes bucinator
      • Piping hornbill, Bycanistes fistulator
      • Silvery-cheeked hornbill, Bycanistes brevis
      • Black-and-white-casqued hornbill, Bycanistes subcylindricus
      • Brown-cheeked hornbill, Bycanistes cylindricus
      • White-thighed hornbill, Bycanistes albotibialis
    • Genus Horizocerus
      • White-crested hornbill, Horizocerus albocristatus
      • Black dwarf hornbill, Horizocerus hartlaubi
    • Genus Tockus
      • Red-billed dwarf hornbill, Tockus camurus
      • Monteiro's hornbill, Tockus monteiri
      • Red-billed hornbill group
        • Northern red-billed hornbill, Tockus erythrorhynchus
        • Damara red-billed hornbill, Tockus damarensis
        • Southern red-billed hornbill, Tockus rufirostris
        • Tanzanian red-billed hornbill, Tockus ruahae
        • Western red-billed hornbill, Tockus kempi
      • Eastern yellow-billed hornbill, Tockus flavirostris
      • Southern yellow-billed hornbill, Tockus leucomelas
      • Jackson's hornbill, Tockus jacksoni
      • Von der Decken's hornbill, Tockus deckeni
      • Crowned hornbill, Tockus alboterminatus
      • Bradfield's hornbill, Tockus bradfieldi
      • African pied hornbill, Tockus fasciatus
      • Hemprich's hornbill, Tockus hemprichii
      • Pale-billed hornbill, Tockus pallidirostris
      • African grey hornbill, Tockus nasutus
    • Genus Ocyceros
      • Malabar grey hornbill, Ocyceros griseus
      • Sri Lanka grey hornbill, Ocyceros gingalensis
      • Indian grey hornbill, Ocyceros biostris
    • Genus Anthracoceros
      • Malabar pied hornbill, Anthracoceros coronatus
      • Oriental pied hornbill, Anthracoceros albirostris
      • Black hornbill, Anthracoceros malayanus
      • Palawan hornbill, Antracoceros marchei
      • Sulu hornbill, Anthracoceros montani
    • Genus Buceros
      • Rhinoceros hornbill, Buceros rhinoceros
      • Great hornbill, Buceros bicornis
      • Rufous hornbill, Buceros hydrocorax
    • Genus Rhinoplax (sometimes included in Buceros)
      • Helmeted hornbill, Rhinoplax vigil
    • Genus Anorrhinus
      • Austen's brown hornbill, Anorrhinus austeni
      • Tickell's brown hornbill, Anorrhinus tickelli
      • Bushy-crested hornbill, Anorrhinus galeritus
    • Genus Penelopides
      • Luzon hornbill, Penelopides manillae
      • Mindoro hornbill, Penelopides mindorensis
      • Visayan hornbill, Penelopides panini
      • Samar hornbill, Penelopides samarensis
      • Mindanao hornbill, Penelopides affinis
    • Genus Berenicornis (sometimes included in Aceros)
      • White-crowned hornbill, Berenicornis comatus
    • Genus Aceros
      • Rufous-necked hornbill, Aceros nipalensis
    • Genus Rhabdotorrhinus
      • Wrinkled hornbill, Rhabdotorrhinus corrugatus
      • Writhed hornbill, Rhabdotorrhinus leucocephalus
      • Rufous-headed hornbill, Rhabdotorrhinus waldeni
      • Sulawesi hornbill, Rhabdotorrhinus exarhatus
    • Genus Rhyticeros (sometimes included in Aceros)
      • Wreathed hornbill, Rhyticeros undulatus
      • Narcondam hornbill, Rhyticeros narcondami
      • Sumba hornbill, Rhyticeros everetti
      • Plain-pouched hornbill, Rhyticeros subruficollis
      • Papuan hornbill, Rhyticeros plicatus
      • Knobbed hornbill, Rhyticeros cassidix
    • Genus Ceratogymna
      • Black-casqued hornbill, Ceratogymna atrata
      • Yellow-casqued hornbill, Ceratogymna elate

References

1. Gill, Frank; Donsker, David, eds. (2019). "Mousebirds, Cuckoo Roller, trogons, hoopoes, hornbills". World Bird List Version 9.2. International Ornithologists' Union. Retrieved 23 July 2019.
2. Johansson, Ulf S.; Ericson, Per G.P. (2003). "Molecular support for a sister group relationship between Pici and Galbulae (Piciformes sensu Wetmore 1960)" (PDF). J. Avian Biol. 34 (2): 185–197. doi:10.1034/j.1600-048X.2003.03103.x. Retrieved 2008-10-30.
3. Yuri, T. et al. (2013) Parsimony and Model-Based Analyses of Indels in Avian Nuclear Genes Reveal Congruent and Incongruent Phylogenetic Signals. Biology, 2(1):419-444. doi:10.3390/biology2010419
4. Jarvis, E.D. et al. (2014) Whole-genome analyses resolve early branches in the tree of life of modern birds. Science, 346(6215):1320-1331.
5. Prum, R.O.; Berv, J.S.; Dornburg, A.; Field, D.J.; Townsend, J.P.; Lemmon, E.M.; Lemmon, A.R. (2015). "A comprehensive phylogeny of birds (Aves) using targeted next-generation DNA sequencing". Nature. 526 (7574): 569–573. Bibcode:2015Natur.526..569P. doi:10.1038/nature15697. PMID 26444237. S2CID 205246158.
6. Woodruff, D. S. & Srikwan, S. 2011. Molecular genetics and the conservation of hornbills in fragmented landscapes. In Poonswad, P. (ed) The Asian Hornbills: Ecology and Conservation. National Center for Genetic Engineering and Biotechnology, Bangkok, pp. 257–264.
7. Fry, C. Hilary (2003). "Wood-hoopoes". In Perrins, Christopher. The Firefly Encyclopedia of Birds. Firefly Books. p. 383. ISBN 1-55297-777-3.
8. Mayr, Gerald (2009). "Paleogene Fossil Birds". In Springer Science & Business Media, 21 April 2009, p. 194. ISBN 978-3-540-89627-2

Further reading

• Gonzalez, J.-C.T.; Sheldon, B.C.; Collar, N.J.; Tobias, J.A. (2013). "A comprehensive molecular phylogeny for the hornbills (Aves: Bucerotidae)". Molecular Phylogenetics and Evolution. 67 (2): 468–483. Bibcode:2013MolPE..67..468G. doi:10.1016/j.ympev.2013.02.012. PMID 23438388.

v t e Genera of trogons, hornbills, kingfishers, woodpeckers and their extinct allies
Kingdom: Animalia Phylum: Chordata Class: Aves Infraclass: Neognathae Clade: Coraciimorphae Clade: Cavitaves
Cavitaves Leptosomiformes Leptosomidae Leptosomus †Plesiocathartes? †Waltonavis Eucavitaves Trogoniformes Trogonidae Apalharpactes Apaloderma †Eotrogon Euptilotis †Foshanornis Harpactes †Masillatrogon †Paratrogon Pharomachrus †Primotrogon Priotelus †Septentrogon Trogon Picocoraciae Bucerotiformes See below ↓ Picodynastornithes Coraciiformes See below ↓ Piciformes See below ↓
Bucerotiformes Bucerotiformes †Laurillardiidae Laurillardia †Messelirrisoridae Messelirrisor Upupi Phoeniculidae †Phirriculus Phoeniculus Rhinopomastus Upupidae Upupa Buceroti Bucorvidae Bucorvus †Euroceros Bucerotidae Aceros Anorrhinus Anthracoceros Berenicornis Buceros Bycanistes Ceratogymna Horizocerus Lophoceros Ocyceros Penelopides Rhabdotorrhinus Rhinoplax Rhyticeros Tockus
Coraciiformes incertae sedis †Cryptornis †Paracoracias †Quasisyndactylus †Eocoraciidae Eocoracias †Geranopteridae Geranopterus †Primobucconidae Primobucco Septencoracias Brachypteraciidae Atelornis Brachypteracias Geobiastes Uratelornis Coraciidae †Chehuenia Coracias Eurystomus †Miocoracias Meropidae Meropogon Merops Nyctyornis Alcedines Alcedinidae Alcedininae Alcedo Ceyx Corythornis Ispidina Cerylinae Ceryle Chloroceryle Megaceryle Halcyoninae Actenoides Caridonax Cittura Clytoceyx Dacelo Halcyon Lacedo Melidora Pelargopsis Syma Tanysiptera Todiramphus Momotidae Aspatha Baryphthengus Electron Eumomota Hylomanes Momotus †Protornis? Todidae †Palaeotodus Todus
Piciformes incertae sedis †"Homalopus" †Pristineanis †Gracilitarsidae Gracilitarsus Neanis †Sylphornithidae Oligosylphe Palaegithalus? Sylphornis Galbuli †Jacamatia Bucconidae Bucco Chelidoptera Hapaloptila Hypnelus Malacoptila Micromonacha Monasa Nonnula Notharchus Nystalus Galbulidae Brachygalba Galbalcyrhynchus Galbula Jacamaralcyon Jacamerops Pici incertae sedis †Rupelramphastoides †Miopiconidae Miopico †Picavidae Picavus Ramphastides Capitonidae Capito †Capitonides Eubucco Lybiidae Buccanodon Cryptolybia Gymnobucco Lybius Pogoniulus Pogonornis Stactolaema Trachyphonus Tricholaema Megalaimidae Caloramphus Psilopogon Ramphastidae Andigena Aulacorhynchus Pteroglossus Ramphastos Selenidera Semnornithidae Semnornis Picides Indicatoridae Indicator Melichneutes Melignomon Prodotiscus Picidae See below ↓
Picidae incertae sedis †Bitumenpicus †Breacopus †Palaeonerpes †Palaeopicus †Piculoides Jynginae Jynx Picumninae Picumnus Sasiinae Sasia Verreauxia Picinae †Australopicus †Palaeonerpes †Pliopicus Nesoctitini Nesoctites Hemicircini Hemicircus Picini Campethera Celeus Chrysophlegma Colaptes Dinopium Dryocopus Gecinulus Geocolaptes Micropternus Meiglyptes Mulleripicus Pardipicus Piculus Picus Campephilini Blythipicus Campephilus Chrysocolaptes Reinwardtipicus Melanerpini Chloropicus Dendrocopos Dendrocoptes Dendropicos Dryobates Leiopicus Leuconotopicus Melanerpes Picoides Sphyrapicus Veniliornis Xiphidiopicus Yungipicus

• Bucerotiformes
• Neognathae
• Bird orders
• Extant Eocene first appearances
• Eocene taxonomic orders
• Oligocene taxonomic orders
• Miocene taxonomic orders
• Pliocene taxonomic orders
• Pleistocene taxonomic orders
• Holocene taxonomic orders
• Taxa named by Max Fürbringer