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A skull of the extinct Gavialis species G. bengawanicus, which lived in the Pleistocene
Gavialis is a genus of crocodylians that includes the living gharialGavialis gangeticus and one known extinct species, Gavialis bengawanicus.G. gangeticus comes from the Indian Subcontinent, while G. bengawanicus is known from Java. Gavialis likely first appeared in the Indian Subcontinent in the Pliocene and dispersed into the Malay Archipelago through a path called the Siva–Malayan route in the Quaternary. Remains attributed to Gavialis have also been found on Sulawesi and Woodlark Island east of the Wallace Line, suggesting a prehistoric lineage of Gavialis was able to traverse marine environments and reach places possibly as far as western Oceania.
The below cladogram of the major extant crocodile groups is based on the latest molecular studies, and shows the gharial's close relationship to the false gharial, and how the gavialids and crocodiles are more closely related than the alligatoroids:
Delfino, M.; De Vos, J. (2010). "A revision of the Dubois crocodylians, Gavialis bengawanicus and Crocodylus ossifragus, from the Pleistocene Homo erectus beds of Java". Journal of Vertebrate Paleontology. 30 (2): 427. Bibcode:2010JVPal..30..427D. doi:10.1080/02724631003617910. S2CID86396515.
Swinton, W. E. (1937). "The crocodile of Maransart (Dollosuchus dixoni )". Mémoires du Musée d'Histoire Naturelle de Belgique. 80: 1–44.
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