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Gnaphalieae
Helichrysum basalticum
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Asterids
Order:	Asterales
Family:	Asteraceae
Subfamily:	Asteroideae
Tribe:	Gnaphalieae Cass. ex Lecoq & Juillet
Genera
See text

The Gnaphalieae, commonly known as paper daisies, are a tribe of flowering plants in the family Asteraceae. It is most closely related to the tribes Anthemideae, Astereae, and Calenduleae.

Characteristics

This group is most diverse in South America, Southern Africa and Australia. There are only a few genera with species native to temperate regions: Anaphalis, Antennaria, Gamochaeta, Helichrysum, Leontopodium (Edelweiss), Phagnalon, Diaperia, and Pseudognaphalium.

The classification of the tribe into subtribes is unclear, with a number of past classifications not being supported by late 20th-century evidence.

Genera

Gnaphalieae genera recognized by the Global Compositae Database as April 2022:

• × Filfia Holub
• Acanthocladium F.Muell.
• Achyrocline (Less.) DC.
• Acomis F.Muell.
• Actinobole Fenzl ex Endl.
• Alatoseta Compton
• Ammobium R.Br.
• Amphiglossa DC.
• Anaphalioides (Benth.) Kirp.
• Anaphalis DC.
• Anaxeton Gaertn.
• Ancistrocarphus A.Gray
• Anderbergia B.Nord.
• Anemocarpa Paul G.Wilson
• Angianthus J.C.Wendl.
• Antennaria Gaertn.
• Apalochlamys Cass.
• Argentipallium Paul G.Wilson
• Argyroglottis Turcz.
• Argyrotegium J.M.Ward & Breitw.
• Arrowsmithia DC.
• Asteridea Lindl.
• Athrixia Ker Gawl.
• Atrichantha Hilliard & B.L.Burtt
• Basedowia E.Pritz.
• Bellida Ewart
• Belloa J.Rémy
• Berroa Beauverd
• Blennospora A.Gray
• Bombycilaena (DC.) Smoljan.
• Bryomorphe Harv.
• Calocephalus R.Br.
• Calomeria Vent.
• Calotesta P.O.Karis
• Cassinia R.Br.
• Castroviejoa Galbany, L.Sáez & Benedí
• Catatia Humbert
• Cephalipterum A.Gray
• Cephalosorus A.Gray
• Chamaepus Wagenitz
• Chevreulia Cass.
• Chiliocephalum Benth.
• Chionolaena DC.
• Chondropyxis D.A.Cooke
• Chryselium Urtubey & S. E. Freire
• Chrysocephalum Walp.
• Chthonocephalus Steetz
• Cladochaeta DC.
• Coronidium Paul G. Wilson
• Craspedia G.Forst.
• Cremnothamnus Puttock
• Cuatrecasasiella H.Rob.
• Dasyanthus Bubani
• Decazesia F.Muell.
• Diaperia Nutt.
• Dicerothamnus Koek.
• Dielitzia P.S.Short
• Disparago Gaertn.
• Dithyrostegia A.Gray
• Dolichothrix Hilliard & B.L.Burtt
• Edmondia Cass.
• Elytropappus Cass.
• Epitriche Turcz.
• Eriochlamys Sond. & F.Muell.
• Erymophyllum Paul G.Wilson
• Euchiton Cass.
• Ewartia Beauverd
• Ewartiothamnus Anderb.
• Facelis Cass.
• Feldstonia P.S.Short
• Filago Loefl.
• Fitzwillia P.S.Short
• Fluminaria N.G.Bergh
• Galeomma Rauschert
• Gamochaeta Wedd.
• Gilberta Turcz.
• Gilruthia Ewart
• Gnaphaliothamnus Kirp.
• Gnaphalium L.
• Gnephosis Cass.
• Gnomophalium Greuter
• Gratwickia F.Muell.
• Haeckeria F.Muell.
• Haegiela P.S.Short & Paul G.Wilson
• Haptotrichion Paul G.Wilson
• Helichrysopsis Kirp.
• Helichrysum Mill.
• Hesperevax (A.Gray) A.Gray
• Humeocline Anderb.
• Hyalochlamys A.Gray
• Hyalosperma Steetz
• Hydroidea P.O.Karis
• Ifloga Cass.
• Ixiolaena Benth.
• Ixodia R.Br.
• Jalcophila M.O.Dillon & Sagást.
• Lachnospermum Willd.
• Langebergia Anderb.
• Laphangium (Hilliard & B.L.Burtt) Tzvelev
• Lasiopogon Cass.
• Lawrencella Lindl.
• Leiocarpa Paul G.Wilson
• Lemooria P.S.Short
• Leontopodium (Pers.) R.Br. ex Cass.
• Lepidostephium Oliv.
• Leptorhynchos Less.
• Leucochrysum (A.Cunn. ex DC.) Paul G.Wilson
• Leucogenes Beauverd
• Leucophyta R.Br.
• Leysera L.
• Logfia Cass.
• Loricaria Wedd.
• Lucilia Cass.
• Metalasia R.Br.
• Mexerion G.L.Nesom
• Micropsis DC.
• Micropus L.
• Millotia Cass.
• Mniodes (A.Gray) Benth. & Hook.f.
• Myriocephalus Benth.
• Myrovernix Koek.
• Neotysonia Dalla Torre & Harms
• Nestlera Spreng.
• Odixia Orchard
• Oedera L.
• Ozothamnus R.Br.
• Paenula Orchard
• Parantennaria Beauverd
• Pentatrichia Klatt
• Petalacte D.Don
• Phaenocoma D.Don
• Phagnalon Cass.
• Pithocarpa Lindl.
• Plecostachys Hilliard & B.L.Burtt
• Podolepis Labill.
• Podotheca Cass.
• Pogonolepis Steetz
• Polycalymma F.Muell. & Sond.
• Pseudognaphalium Kirp.
• Psilocarphus Nutt.
• Pterochaeta Steetz
• Pterygopappus Hook.f.
• Pycnosorus Benth.
• Quinetia Cass.
• Quinqueremulus Paul G.Wilson
• Rachelia J.M.Ward & Breitw.
• Raoulia Hook.f. ex Raoul
• Raouliopsis S.F.Blake
• Rhetinocarpha Paul G.Wilson & M.A.Wilson
• Rhodanthe Lindl.
• Rhynchopsidium DC.
• Rutidosis DC.
• Schoenia Steetz
• Scyphocoronis A.Gray
• Seriphium L.
• Siemssenia Steetz
• Siloxerus Labill.
• Sinoleontopodium Y.L.Chen
• Sondottia P.S.Short
• Stenocline DC.
• Stenophalium Anderb.
• Stoebe L.
• Stuartina Sond.
• Stylocline Nutt.
• Swammerdamia DC.
• Syncarpha DC.
• Syncephalum DC.
• Taplinia Lander
• Tenrhynea Hilliard & B.L.Burtt
• Thiseltonia Hemsl.
• Tietkensia P.S.Short
• Toxanthes Turcz.
• Trichanthodium Sond. & F.Muell.
• Triptilodiscus Turcz.
• Troglophyton Hilliard & B.L.Burtt
• Vellereophyton Hilliard & B.L.Burtt
• Waitzia J.C.Wendl.
• Xerochrysum Tzvelev

Other genera

• Callilepis (may not belong in this tribe)
• Libinhania
• Omalotheca
• Oxylaena
• Planea

References

1. Nie, Ze‐Long; Funk, Vicki A.; Meng, Ying; Deng, Tao; Sun, Hang; Wen, Jun (March 2016). "Recent assembly of the global herbaceous flora: evidence from the paper daisies (Asteraceae: Gnaphalieae)". New Phytologist. 209 (4): 1795–1806. doi:10.1111/nph.13740. ISSN 0028-646X.
2. Panero, JL; VA Funk (2002-12-30). "Toward a phylogenetic subfamilial classification for the Compositae (Asteraceae)" (PDF). Proceedings of the Biological Society of Washington. 115 (4). Biological Society of Washington: 909–922. Retrieved 2010-08-28.
3. ^ Randall J. Bayer; Christopher F. Puttock; Scot A. Kelchner (2000). "Phylogeny of South African Gnaphalieae (Asteraceae) based on two noncoding chloroplast sequences". American Journal of Botany. 87 (2): 259–272. doi:10.2307/2656914. JSTOR 2656914. PMID 10675314.
4. "Gnaphalieae Cass. ex Lecoq & Juill". Global Compositae Database. Compositae Working Group (CWG). 2022. Retrieved 2022-04-21.
5. "Gnaphaliinae Dumort". Global Compositae Database. Compositae Working Group (CWG). 2022. Retrieved 2022-04-21.
6. "Relhaniinae Less". Global Compositae Database. Compositae Working Group (CWG). 2022. Archived from the original on 2022-04-21. Retrieved 2022-04-21.
7. Kilian, Norbert; Galbany-Casals, Mercè; Sommerer, Ronny; Oberprieler, Christoph; Smissen, Rob; Miller, Antony; Rabe, Katharina (2017). "Systematics of Libinhania, a new endemic genus of Gnaphalieae (Asteraceae) from the Socotra archipelago (Yemen), inferred from plastid, low-copy nuclear and nuclear ribosomal DNA loci". Botanical Journal of the Linnean Society. 183 (3): 373–412. doi:10.1093/botlinnean/bow013.

External links

• Data related to Gnaphalieae at Wikispecies
• Media related to Gnaphalieae at Wikimedia Commons
• UniProt. "Tribe Gnaphalieae". Retrieved 2008-05-16.

• Gnaphalieae
• Asteraceae tribes