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Shuotheriidae Temporal range: Middle-Late Jurassic PreꞒ Ꞓ O S D C P T J K Pg N
Life restoration of Pseudotribos
Lower jaw of Shuotherium dongi
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Synapsida
Clade:	Therapsida
Clade:	Cynodontia
Clade:	Mammaliaformes
Family:	†Shuotheriidae Chow and Rich, 1982
Genera
†Feredocodon †Pseudotribos †Shuotherium †Itatodon? †Paritatodon?

Shuotheriidae is a small family of Jurassic mammaliaforms whose remains are found in China, Great Britain and possibly Russia. They have been proposed to be close relatives of Australosphenida (which often controversially includes monotremes), together forming the clade Yinotheria. However, some studies suggest shuotheres are closer to therians than to monotremes, or that australosphenidans and therians are more closely related to each other than either are to shuotheres, with a 2024 study suggesting that shuotheriids were closely related to Docodonta outside of the Mammalia crown group.

References

1. Mao, F.; Li, Z.; Wang, Z.; Zhang, C.; Rich, T.; Vickers-Rich, P.; Meng, J. (2024). "Jurassic shuotheriids show earliest dental diversification of mammaliaforms". Nature. doi:10.1038/s41586-024-07258-7.
2. ^ Wang, Y.-Q. and Li, C.-K. 2016. Reconsideration of the systematic position of the Middle Jurassic mammaliaforms Itatodon and Paritatodon. Palaeontologia Polonica 67, 249–256.
3. Rougier, Guillermo W.; Martinelli, Agustín G.; Forasiepi, Analía M.; Novacek, Michael J., New Jurassic mammals from Patagonia, Argentina : a reappraisal of australosphenidan morphology and interrelationships; American Museum Novitates, no. 3566, 2007
4. Tom Rich, Patricia Vickers Rich, Palaeobiogeography of Mesozoic Mammals – Revisited, Article · January 2012 doi:10.1007/978-90-481-3428-1_32
5. Averianov, A. O.; Lopatin, A. V. (July 2014). "On the phylogenetic position of monotremes (Mammalia, Monotremata)". Paleontological Journal. 48 (4): 426–446. doi:10.1134/s0031030114040042. ISSN 0031-0301. S2CID 84039223 – via Scientific Index.
6. Ramírez-Chaves, Héctor E.; Weisbecker, Vera; Wroe, Stephen; Phillips, Matthew J. (December 2016). "Resolving the evolution of the mammalian middle ear using Bayesian inference". Frontiers in Zoology. 13 (1): 39. doi:10.1186/s12983-016-0171-z. ISSN 1742-9994. PMC 4997658. PMID 27563341.
7. Mao, Fangyuan; Li, Zhiyu; Wang, Zhili; Zhang, Chi; Rich, Thomas; Vickers-Rich, Patricia; Meng, Jin (2024-04-03). "Jurassic shuotheriids show earliest dental diversification of mammaliaforms". Nature. doi:10.1038/s41586-024-07258-7. ISSN 0028-0836.

Further reading

• Zofia Kielan-Jaworowska, Richard L. Cifelli, and Zhe-Xi Luo, Mammals from the Age of Dinosaurs: Origins, Evolution, and Structure (New York: Columbia University Press, 2004), 214–215.

v t e Cynodontia
Kingdom: Animalia Phylum: Chordata Clade: Synapsida Clade: Therapsida Clade: Eutheriodontia Synapsida see Synapsida Cynodontia see below↓
Cynodontia Cynodontia †Dvinia †Nanocynodon †Madysaurus †Uralocynodon †Charassognathidae Abdalodon Charassognathus Nshimbodon †Procynosuchidae Procynosuchus Sludica Epicynodontia †Bolotridon †Cynosaurus †Nanictosaurus †Nythosaurus †Platycraniellus †Thrinaxodon †Vetusodon †Galesauridae Galesaurus Progalesaurus Eucynodontia †Kataigidodon †Cynognathia see Cynognathia Probainognathia see below↓
Probainognathia Probainognathia †Agudotherium †Candelariodon †Charruodon †Lumkuia †Protheriodon †Tessellatia †Chiniquodontidae Aleodon Chiniquodon Cromptodon? Riojanodon †Probainognathidae Bonacynodon Probainognathus †Ecteniniidae Diegocanis Ecteninion Trucidocynodon Prozostrodontia †Alemoatherium †Deccanodon †Dromatherium †Inditherium †Microconodon †Polonodon †Pseudotriconodon †Rewaconodon †Santacruzgnathus †Therioherpeton †Tricuspes †Prozostrodontidae Prozostrodon Pseudotherium †Tritheledontidae Chaliminia Diarthrognathus Elliotherium Irajatherium Pachygenelus Riograndia? Tritheledon Mammaliamorpha †Adelobasileus †Botucaraitherium †Brasilodon †Sinoconodon †Tikiodon †Tritylodontidae Bienotherium Bienotheroides Bocatherium Dianzhongia Dinnebitodon Fossiomanus Kayentatherium Lufengia Montirictus Nuurtherium Oligokyphus Polistodon Shartegodon Stereognathus Tritylodon Xenocretosuchus Yuanotherium Yunnanodon Mammaliaformes see below↓
Mammaliaformes Mammaliaformes †Bocaconodon †Brachyzostrodon †Bridetherium †Cherwellia †Cifellilestes †Delsatia †Dianoconodon †Dinnetherium †Dyskritodon †Eozostrodon †Erythrotherium †Gondwanadon †Hadrocodium †Hallautherium †Helvetiodon †Indotherium †Indozostrodon †Megazostrodon †Morganucodon †Paceyodon †Paikasigudodon †Purbeckodon †Rosierodon †Sinoconodon? †Storchodon †Stylidens †Wareolestes †Woutersia †Kuehneotheriidae Fluctuodon Kotatherium Kuehneon Kuehneotherium †Shuotheriidae Feredocodon Itatodon? Paritatodon? Pseudotribos Shuotherium †Docodonta Agilodocodon Borealestes Castorocauda Cyrtlatherium Dobunnodon Docodon Docofossor Dsungarodon Ergetiis Gondtherium? Haldanodon Hutegotherium Itatodon? Khorotherium Krusatodon Microdocodon Paritatodon? Peraiocynodon Sibirotherium Simpsonodon Tashkumyrodon Tegotherium †Haramiyida Avashishta? Hypsiprymnopsis? Haramiyavia Kalaallitkigun Kollikodon? Mojo? Theroteinus Thomasia Euharamiyida Allostaffia? Arboroharamiya Baidabatyr? Butlerodon Cryoharamiya Kermackodon? Maiopatagium Megaconus? Qishou Sharypovoia Shenshou Sineleutherus Vilevolodon Woodeatonia Xianshou Gondwanatheria? Hahnodontidae? Cifelliodon Denisodon Hahnodon Crown-Mammalia see Mammalia
See also Incertae sedis †Cistecynodon †Eoraetia †Gaumia †"Hahnia" †Kunminia †Lepagia †Meurthodon †Mitredon †Panchetocynodon †Redondagnathus †Saurodesmus? Paraphyletic / Polyphyletic groups Dromatheriidae Morganucodonta Symmetrodonta Triconodonta
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• Shuotheriidae
• Mammaliaformes
• Prehistoric therapsid families
• Middle Jurassic first appearances
• Late Jurassic extinctions
• Taxa named by Zhou Mingzhen
• Taxa named by Tom Rich
• Cynodont stubs