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Vjushkovisaurus Temporal range: Middle Triassic, Anisian PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	Archosauromorpha
Clade:	Archosauriformes
Genus:	†Vjushkovisaurus Ochev, 1982
Type species
†Vjushkovisaurus berdjanensis Ochev, 1982

Vjushkovisaurus is an extinct genus of Middle Triassic archosauriform. It is known from the Anisian-aged Donguz Gorizont in Sol-Iletsk, Orenburg Oblast, Russia. The genus was named in 1982, with the type species being V. berdjanensis. Material has been collected in the Berdyanka II locality from a fossil assemblage called the Eryosuchus Fauna along the Berdyanka River, specifically in a sand-carbonate concretion in the upper part of the main river channel. Vjushkovisaurus is known only from the holotype PIN 2865/62 (formerly SGU 104/3871), a partial postcranial skeleton which consists of 12 presacral vertebrae, left humerus, ribs, a fragment of the coracoid and a fragment of the fibula.

Description

Most rauisuchids have a crest on the ilium called the supra-acetabular process that overlies the head of the femur, allowing them to have a "pillar-erect" stance. As an early rauisuchid Vjushkovisaurus lacks this crest, but it does have a small thickening on the surface of the ilium. The humerus differs from other better known rauisuchids in that it is shorter and broader. However, it has been suggested that the only known humerus of Vjushkovisaurus may actually belong to an aetosaur.

More recently, Nesbitt (2009) argued that Vjushkovisaurus represent most probably a valid non-archosaurian archosauriform. According to his analysis, the original diagnosis of Vjushkovisaurus by Ochev (1982) consisted of some of the plesiomorphies for Archosauriformes and characters with wide distributions throughout archosauriforms. For example, elongated anterior cervical vertebrae are present in the poposauroids, Yarasuchus, and in the non-archosaurian archosauriforms Proterosuchus and Guchengosuchus. Other features which are typically thought to be associated with skeletal pneumaticity, are also present in Guchengosuchus, Erythrosuchus, and in several archosaurs. Additionally, the shape and placement of the deltopectoral crests of the humerus of Vjushkovisaurus are typical of basal archosauriforms. The ectepicondylar groove on the humerus reported by Ochev is present in Prolacerta, Proterosuchus, phytosaurs, aetosaurs and several paracrocodylomorphs. Gower and Sennikov (2000) reported that the posterior cervical vertebrae bear well-defined facets for three-headed ribs. According to Nesbitt (2009), this trait is absent in Archosauria with the possible exception of Arizonasaurus and Poposaurus, but was used as a synapomorphy of Erythrosuchia in Parrish (1992) phylogenetic analysis.

Paleobiology

Other vertebrates from the Berdyanka II locality that lived alongside Vjushkovisaurus include capitosauroid temnospondyls and the therapsid Nothogomphodon, one of the last therocephalians. Vjushkovisaurus was likely a top predator in its environment along other large carnivores including Dongusuchus and Erythrosuchus.

References

1. Ochev, V.G. (1982). "Psevdosukhii iz Srednego Triasa Yuzhnogo Priural'ya". Paleontologicheskii Zhurnal. 1982: 96–102.
2. ^ Gower, D.J.; Sennikov, A.G. (1997). "Sarmatosuchus and the early history of the Archosauria". Journal of Vertebrate Paleontology. 17 (1): 60–73. doi:10.1080/02724634.1997.10010954.
3. ^ Sterling J. Nesbitt (2009). "The early evolution of archosaurs: Relationships and the origin of major clades". Columbia University (Open Access Dissertation): 1–632.
4. Bonaparte, J.F. (1984). "Locomotion in rauisuchid thecodonts". Journal of Vertebrate Paleontology. 3 (4): 210–218. doi:10.1080/02724634.1984.10011976.
5. Benton, M.J.; Clark, J. (1988). "Archosaur phylogeny and the relationships of the Crocodylia". In Benton, M.J. (ed.). The Phylogeny and Classification of the Tetrapods. 1. Amphibians, Reptiles, Birds (PDF). Systematics Association. Vol. 35A. Oxford: Clarendon Press. pp. 289–332. Archived from the original (PDF) on July 15, 2007.
6. Gower, D.J.; Sennikov, A.G. (2003). "Early archosaurs from Russia". In Benton, M.J.; Shishkin, M.A.; Unwin, D.M. (eds.). The Age of Dinosaurs in Russia and Mongolia. Cambridge: Cambridge University Press. pp. 140–159.
7. Juul, L. (1994). "The phylogeny of basal archosaurs". Palaeontologica Africana. 31: 1–38.
8. Sennikov, A.G. (1996). "Evolution of the Permian and Triassic tetrapod communities of Eastern Europe". Palaeogeography, Palaeoclimatology, Palaeoecology. 120 (3): 331–351. Bibcode:1996PPP...120..331S. doi:10.1016/0031-0182(95)00041-0.

External links

• Vjushkovisaurus in the Paleobiology Database

v t e Archosauromorpha
Kingdom: Animalia Phylum: Chordata Class: Sauropsida Clade: Diapsida Clade: Sauria Clade: Archelosauria Sauropsida see Sauropsida Archosauromorpha see below↓
Archosauromorpha Archosauromorpha †Aenigmastropheus †Czatkowiella †Elachistosuchus? †Elessaurus †Malutinisuchus †Mecistotrachelos †Prolacertoides †Protorosaurus †Rhombopholis †Vritramimosaurus †Tanysauria Jesairosaurus? Trachelosauridae Austronaga Dinocephalosaurus Fuyuansaurus? Gracilicollum? Protanystropheus Pectodens Sclerostropheus? Trachelosaurus Sharovipterygidae? Ozimek Sharovipteryx Tanystropheidae Amotosaurus Augustaburiania Cosesaurus Fuyuansaurus? Gracilicollum? Gwyneddosaurus Langobardisaurus Luxisaurus Macrocnemus Raibliania Sclerostropheus? Tanystropheus Tanytrachelos Sharovipterygidae? Crocopoda †Boreopricea †Cryptovaranoides? †Tasmaniosaurus †Teyujagua †Allokotosauria see Allokotosauria †Rhynchosauria see Rhynchosauria †Prolacertidae? Kadimakara Prolacerta Archosauriformes see below↓
Archosauriformes Archosauriformes †Antarctanax †Cuyosuchus †Eorasaurus? †Kalisuchus †Koilamasuchus †Sarmatosuchus †Syntomiprosopus †Proterosuchidae Archosaurus Chasmatosuchus? Gamosaurus Jaikosuchus Proterosuchus Samsarasuchus Tsylmosuchus? Vonhuenia? †Protopyknosia Kranosaura Triopticus †Erythrosuchidae Bharitalasuchus Chalishevia Erythrosuchus Fugusuchus? Garjainia Guchengosuchus Shansisuchus Uralosaurus Eucrocopoda †Asperoris †Dorosuchus †Heteropelta †Litorosuchus †Marcianosuchus †Polymorphodon †Vancleavea †Euparkeriidae Euparkeria Halazhaisuchus Osmolskina †Proterochampsia see Proterochampsia Crurotarsi †Phytosauria see Phytosauria Archosauria Incertae sedis †Sikannisuchus? †Smok Avemetatarsalia see Avemetatarsalia includes birds Pseudosuchia see Pseudosuchia includes crocodilians
Related topics Tooth taxa †Crosbysaurus †Lucianosaurus †Tecovasaurus †Palaeosaurus †Protecovasaurus †Uatchitodon †Zanclodon Nomina dubia †Albisaurus †Ankistrodon †Avipes †Blomosuchus †Collilongus †Exilisuchus †Fenhosuchus †Megacnemus †Microcnemus †Vjushkovisaurus †Wangisuchus Paraphyletic groups Proterosuchia Protorosauria Thecodontia Possible members Choristodera Drepanosauromorpha Kuehneosauridae Ichthyosauromorpha Sauropterygia Thalattosauria
Category

v t e Pseudosuchia
Kingdom: Animalia Phylum: Chordata Class: Sauropsida Clade: Archosauria Sauropsida see Sauropsida Archosauromorpha see Archosauromorpha Pseudosuchia see below↓
Pseudosuchia Pseudosuchia †Dasygnathoides †Nundasuchus †Phytosauria? †Ornithosuchidae Aenigmaspina? Dynamosuchus Ornithosuchus Riojasuchus Venaticosuchus Suchia †Heteropelta? †Koilamasuchus? †Mandasuchus †Nundasuchus? †Ticinosuchus †Erpetosuchidae Archeopelta? Dyoplax? Erpetosuchus Pagosvenator Parringtonia Tarjadia †Aetosauriformes Acaenasuchus Euscolosuchus Revueltosaurus Aetosauria see Aetosauria †Gracilisuchidae Gracilisuchus Maehary? Parvosuchus Turfanosuchus Yonghesuchus Paracrocodylomorpha see below↓
Paracrocodylomorpha Paracrocodylomorpha †Mambawakale †Poposauroidea Benggwigwishingasuchus Lotosaurus Schultzsuchus? Qianosuchus Ctenosauriscidae Arizonasaurus Bromsgroveia Bystrowisuchus Ctenosauriscus Hypselorhachis Tsylmosuchus? Xilousuchus Poposauridae Dolichobrachium Poposaurus Shuvosauridae Effigia Shuvosaurus Sillosuchus Loricata †Apatosuchus †Arganasuchus †Batrachotomus †Dagasuchus †Decuriasuchus †Etjosuchus †Fasolasuchus †Heptasuchus †Heteropelta? †Incertovenator? †Luperosuchus †Mandasuchus? †Prestosuchus †Procerosuchus †Saurosuchus †Stagonosuchus †Youngosuchus †Rauisuchidae Arganasuchus? Energosuchus? Jushatyria? Polonosuchus Postosuchus Procerosuchus? Rauisuchus Scolotosuchus Scythosuchus? Teratosaurus Tikisuchus? Vivaron Vjushkovisaurus? Vytshegdosuchus? Crocodylomorpha see below↓
Crocodylomorpha Crocodylomorpha †Carnufex †Dibothrosuchus †Dromicosuchus †Hesperosuchus †Heteropelta? †Junggarsuchus †Kayentasuchus †Parrishia †Phyllodontosuchus †Pseudhesperosuchus †Redondavenator †Sphenosuchus †Syntomiprosopus? †Trialestes †Saltoposuchidae Litargosuchus Saltoposuchus Terrestrisuchus †Hallopodidae Almadasuchus Hallopus Macelognathus †Thalattosuchia see Thalattosuchia Crocodyliformes †Entradasuchus †Eopneumatosuchus †Kemkemia †Microchampsa †Platyognathus †Stegomosuchus †Tagarosuchus †Protosuchidae Coloradisuchus Dianosuchus Edentosuchus Erythrochampsa Hemiprotosuchus Hoplosuchus Orthosuchus? Pedeticosaurus? Protosuchus †Notochampsidae Notochampsa Orthosuchus? †Gobiosuchidae Cassissuchus Gobiosuchus Lisboasaurus? Zaraasuchus Mesoeucrocodylia †Dianchungosaurus †Hsisosuchus †Induszalim †Lusitanisuchus †Neuquensuchus †Pachycheilosuchus? †Titanochampsa †Thalattosuchia? †Shartegosuchoidea Hadongsuchus Shantungosuchus Sichuanosuchus Zosuchus Shartegosuchidae Adzhosuchus Fruitachampsa Kyasuchus Nominosuchus Shartegosuchus Metasuchia †Ziphosuchia see Ziphosuchia includes notosuchians Neosuchia see Neosuchia includes crocodilians
See also: Rauisuchia Prestosuchidae Bathyotica Sphenosuchia Protosuchia Mesosuchia Category

• Archosauriforms
• Prehistoric reptile genera
• Anisian genera
• Middle Triassic reptiles of Europe
• Triassic Russia
• Fossils of Russia
• Fossil taxa described in 1982