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{{Short description|Large cat native to the Americas}}
{{alternateuses}}
{{About||the car manufacturer|Jaguar Cars|other uses}}
{{Taxobox
{{Featured article}}
| color = pink
{{Pp|small=yes}}
{{pp-move}}
{{Use American English|date=May 2024}}
{{Use dmy dates|date=May 2024}}
{{Speciesbox
| name = Jaguar | name = Jaguar
| fossil_range = {{longitem|style=line-height:1.25em|{{nowrap|] – ]}} {{nowrap|(~850,000–0 ])<ref name=SK1993>{{cite book |author=Seymour, K.|editor1=Martin, R. A. |editor2=Barnosky, A. D. |year=1993 |chapter=Size change in North American Quaternary jaguars |title=Morphological change in Quaternary mammals of North America |publisher=Cambridge University Press |pages=343–372 |doi=10.1017/CBO9780511565052.014|isbn=978-0-521-40450-1 }}</ref>}}}}
| image = Standing jaguar.jpg <!--Please do not change this image. See Talkpage for two (Feb 2021 & Jul 2021) consensus-based sections supporting this (original FA) image. Thanks!-->
| image_upright = 1.2
| status = NT | status = NT
| status_system = IUCN3.1
| trend = down
| status_ref = <ref name=iucn>{{cite iucn |title=''Panthera onca'' |author1=Quigley, H. |author2=Foster, R. |author3=Petracca, L. |author4=Payan, E. |author5=Salom, R. |author6=Harmsen, B. |year=2017 |errata=2018 |name-list-style=amp |page=e.T15953A123791436 |doi=10.2305/IUCN.UK.2017-3.RLTS.T15953A50658693.en |access-date=15 January 2022}}</ref>
| image = Panthera onca smithsonian.jpg
| image_width = 200px | status2 = CITES_A1
| status2_system = CITES
| regnum = ]ia
| status2_ref = <ref name=iucn/>
| phylum = ]
| taxon = Panthera onca
| classis = ]ia
| authority = (], ])
| ordo = ]
| subdivision_ranks = Subspecies
| familia = ]
| subdivision =
| genus = '']''
| species = '''''P. onca''''' *''Panthera onca onca''
| binomial = ''Panthera onca'' *{{extinct}}'']''
*{{extinct}}'']''
| binomial_authority = (], ])
| range_map = Panthera onca distribution.svg
| range_map_caption = {{color box|red}} Current range
{{color box|pink}} Former range
| synonyms = {{Collapsible list
| title = {{Clear}}
| bullets = yes
| ''Felis augustus'' {{small|(Leidy, 1872)}}
| ''Felis listai'' {{small|(Roth, 1899)}}
| ''Felis onca'' {{small|Linnaeus, 1758}}
| ''Felis onca'' subsp. ''boliviensis'' {{small|Nelson & Goldman, 1933}}
| ''Felis onca'' subsp. ''coxi'' {{small|Nelson & Goldman, 1933}}
| ''Felis onca'' subsp. ''ucayalae'' {{small|Nelson & Goldman, 1933}}
| ''Felis veronis'' {{small|Hay, 1919}}
| ''Iemish listai'' {{small|(Roth, 1899)}}
| ''Panthera augusta'' {{small|(Leidy, 1872)}}
| ''Panthera onca'' subsp. ''augusta'' {{small|(Leidy, 1872)}}
| ''Uncia augusta'' {{small|(Leidy, 1872)}}
}}
| synonyms_ref = <ref name=MSW3>{{MSW3 Carnivora |id=14000240 |pages=546–547 |heading=Species ''Panthera onca''}}</ref>
}} }}


The '''jaguar''' ('''''Panthera onca''''') is a large ] species and the only ] member of the genus '']'' that is native to the ]. With a body length of up to {{cvt|1.85|m|ftin}} and a weight of up to {{cvt|158|kg}}, it is the biggest cat species in the Americas and the ] in the world. Its distinctively marked ] features pale yellow to tan colored fur covered by spots that transition to ] on the sides, although a ] black coat appears in some individuals. The jaguar's powerful bite allows it to pierce the ]s of ]s and ]s, and to employ an unusual killing method: it bites directly through the skull of ]ian ] between the ears to deliver a fatal blow to the brain.
The '''jaguar''' (''Panthera onca'') are ]s of the ] family and one of four "]s" in the '']'' ]. It is closely related to the ], ], and ] of the ], and is the largest and most powerful feline in the ]. It is the third largest feline after the lion and tiger.


The modern jaguar's ancestors probably entered the Americas from ] during the ] via the ] that once spanned the ]. Today, the jaguar's range extends from the ] across ] and much of ], the ] and south to ] and northern ]. It inhabits a variety of forested and open terrains, but its preferred ] is ], ]s and wooded regions. It is adept at swimming and is largely a solitary, opportunistic, stalk-and-ambush ]. As a ], it plays an important role in stabilizing ]s and in regulating prey populations.
==Physical characteristics==
To some, jaguars look very much like ], but they are sturdier and heavier. The easiest way to distinguish a jaguar from a leopard, besides the jaguar’s much more powerful build, is by the ]. The rosettes on a jaguar’s coat are larger, fewer in number, and usually darker with thicker lines that enclose smaller spots. The head of the jaguar is rounder and it has shorter, stockier limbs. Because of this the jaguar is sometimes referred to as the “bulldog” of the cat world. The Jaguar, in a recent ] special titled “In Search of the Jaguar,” was named pound for pound the most powerful cat in the world.


The jaguar is threatened by ], ], ] for trade with its body parts and killings in ] situations, particularly with ]ers in Central and ]. It has been listed as ] on the ] since 2002. The wild population is thought to have declined since the late 1990s. Priority areas for jaguar ] comprise 51 Jaguar Conservation Units (JCUs), defined as large areas inhabited by at least 50 breeding jaguars. The JCUs are located in 36 geographic regions ranging from Mexico to Argentina.
Jaguars vary from 5.3 to 6 feet (1.62 to 1.83 m) in length, including a 30 in (0.76 m) tail. They stand around 67 to 76 cm (27 to 30 inches) tall at the shoulder and weigh between 56 and 96 kg (124 and 211 ]) with larger individuals, recorded by scientists, weighing between 131 and 151 kg (288 to 333 lb). Females are typically twenty percent smaller than males. Jaguars in southern Mexico and Central America are typically smaller--56 kg and 40 kg (123 lb and 90 lb) for males and females respectively. .


The jaguar has featured prominently in the mythology of ], including those of the ] and ]s.
Jaguars are powerful animals for their size."They are powerful enough to drag an 800-pound bull 25 feet in its jaws and pulverize the heaviest bones."
A predator needs to be strong and fast enough to catch its prey. Jaguars hunt wild animals in the range of 300 kg and below in dense jungle; their short and sturdy physique is thus an adaptation to hunt those animals in that environment.


==Habitat== ==Etymology==
The word "jaguar" is possibly derived from the ] word {{lang|und|yaguara}} meaning 'wild beast that overcomes its prey at a bound'.<ref name=Guggisberg1975>{{Cite book |last=Guggisberg |first=C. A. W. |title=Wild Cats of the World |year=1975 |publisher=Taplinger Publishing |location=New York |isbn=978-0-8008-8324-9 |chapter=Jaguar ''Panthera onca'' (Linnaeus, 1758) |pages=247–265}}</ref><ref name=Seymour/> In North America, the word is pronounced disyllabic {{IPAc-en|ˈ|dʒ|æ|g|w|ɑːr}}, while in British English, it is pronounced with three syllables {{IPAc-en|ˈ|dʒ|æ|g|juː|ər}}.<ref>{{cite book |author=Qualls, E. J. |year=2012 |title=The Qualls Concise English Grammar |publisher=Danaan Press |isbn=9781890000097 |chapter=The dialects of English |chapter-url=https://books.google.com/books?id=tjrd5UnFT2gC&pg=PT29}}</ref><ref>{{cite web |title=Jaguar |url=https://dictionary.cambridge.org/us/pronunciation/english/jaguar |website=] |access-date=16 November 2021 |archive-date=12 November 2020 |archive-url=https://web.archive.org/web/20201112012608/https://dictionary.cambridge.org/us/pronunciation/english/jaguar |url-status=live}}</ref> Because that word also applies to other animals, ] call it {{lang|gyn|jaguareté}}, with the added sufix ''eté'', meaning "true beast".<ref>{{cite book |author=Labat, J.B. |author-link=Jean-Baptiste Labat |year=1731 |chapter=Once, espèce de Tigre |title=Voyage du chevalier Des Marchais en Guinée, isles voisines, et à Cayenne, fait en 1725, 1726 & 1727 |location=Amsterdam |publisher=La Compagnie |volume=III |page=285 |chapter-url=https://books.google.com/books?id=dfdWAAAAcAAJ&pg=RA6-PA5 |access-date=13 August 2020 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021340/https://books.google.com/books?id=dfdWAAAAcAAJ&pg=RA6-PA5 |url-status=live}}</ref>
]
"Onca" is derived from the Portuguese name {{lang|pt|onça}} for a spotted cat that is larger than a ]; cf. ].<ref>{{cite book |author=Ray, J. |author-link=John Ray |year=1693 |chapter=Pardus an Lynx brasiliensis, ''Jaguara'' |title=Synopsis Methodica Animalium Quadrupedum et Serpentini Generis. Vulgarium Notas Characteristicas, Rariorum Descriptiones integras exhibens |publisher=S. Smith & B. Walford |location=London |chapter-url=https://archive.org/details/synopsismethodic00rayj/page/168/mode/1up |page=168}}</ref> The word "panther" is derived from ] {{lang|la|panthēra}}, itself from the ] {{lang|grc|πάνθηρ}} ({{transliteration|grc|pánthēr}}).<ref>{{cite book |author1=Liddell, H. G. |author2=Scott, R. |year=1940 |name-list-style=amp |script-chapter=el:πάνθηρ |title=A Greek-English Lexicon |edition=Revised and augmented |location=Oxford |publisher=Clarendon Press |url=https://www.perseus.tufts.edu/cgi-bin/ptext?doc=Perseus%3Atext%3A1999.04.0057%3Aentry%3D%2377441 |access-date=20 February 2021 |archive-date=11 April 2008 |archive-url=https://web.archive.org/web/20080411203109/http://www.perseus.tufts.edu/cgi-bin/ptext?doc=Perseus%3Atext%3A1999.04.0057%3Aentry%3D%2377441 |url-status=live}}</ref>
The jaguar's habitat ranges from the rain forests of ] and ] to marshy and even desert terrain in ], but they are rarely seen in mountainous regions. The jaguar's wide range means that it should not be in danger of extinction in the foreseeable future. The species has declined in numbers in some areas due to habitat loss, especially in rain forests and grassland turned into cropland and hunting for their pelts.


== Taxonomy and evolution ==
Known for their strong swimming abilities, the jaguar is one of the few cats besides tigers that enjoy water. They often prefer to live by rivers, swamps, and in dense forest with thick cover for stalking ]. Jaguars, on rare occasions, are seen as far north as the southwestern ], particularly in ], ] and ]. In the early 1900s, the jaguars' range actually extended as far north as Southern ] and western ]. As recently as 2004, wildlife officials in ] have photographed and documented jaguars in the southern parts of the state. Presently it is unclear whether recent sightings indicate whether there is a permanent population developing in the Southwest or that these cats are simply transients straying over the border from ]. However, jaguars are a protected species in the ] under the ] and are considered nongame, therefore making it illegal to shoot a jaguar for its pelt. Fossils of jaguars from as far north as ] confirm these cats inhabited much of the Southern United States during prehistoric times. These prehistoric jaguars were significantly larger than the jaguars of today.
===Taxonomy===
In 1758, ] described the jaguar in his work '']'' and gave it the ] ''Felis onca''.<ref name=Linn1758>{{cite book |author=Linnaeus, C. |year=1758 |chapter=''Felis onca'' |title=Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis |edition=Decima, reformata |language=la |location=Holmiae |publisher=Laurentius Salvius |volume=I |page=42 |chapter-url=https://archive.org/stream/mobot31753000798865#page/41/mode/2up}}</ref>


In the 19th and 20th centuries, several jaguar ]s formed the basis for descriptions of ].<ref name=MSW3/> In 1939, ] recognized eight subspecies based on the geographic origins and skull ] of these specimens.<ref name=Pocock1939>{{cite journal |last=Pocock |first=R. I. |year=1939 |title=The races of jaguar (''Panthera onca'') |journal=Novitates Zoologicae |volume=41 |pages=406–422 |url=https://archive.org/details/cbarchive_123320_theracesofjaguarpantheraonca9999}}</ref>
==Taxonomy==
Pocock did not have access to sufficient ]s to critically evaluate their subspecific status but expressed doubt about the status of several. Later consideration of his work suggested only three subspecies should be recognized. The description of ''P. o. palustris'' was based on a ] skull.<ref name=Seymour>{{cite journal |last=Seymour |first=K. L. |year=1989 |title=''Panthera onca'' |journal=Mammalian Species |issue=340 |pages=1–9 |url=http://www.science.smith.edu/departments/Biology/VHAYSSEN/msi/pdf/i0076-3519-340-01-0001.pdf |url-status=dead |archive-url=https://web.archive.org/web/20100620231016/http://www.science.smith.edu/departments/Biology/VHAYSSEN/msi/pdf/i0076-3519-340-01-0001.pdf |archive-date=20 June 2010 |doi=10.2307/3504096 |jstor=3504096|s2cid=253932256 }}</ref>
===Species===
Todays recognised species:
*''Panthera onca''


By 2005, nine subspecies were considered to be ] taxa:<ref name=MSW3 />
There are also prehistoric (sub)species:
*''P. o. onca'' {{small|(Linnaeus, 1758)}} was a jaguar from Brazil.<ref name=Linn1758 />
*''Panthera toscana'' (])
*''P. o. peruviana'' {{small|(], 1843)}} was a jaguar skull from Peru.<ref>{{cite book |author=Blainville, H. M. D. de |year=1843 |chapter=''F. leo nubicus'' |title=Ostéographie ou description iconographique comparée du squelette et du système dentaire des mammifères récents et fossils pour servir de base à la zoologie et la géologie |language=fr |location=Paris |publisher=J. B. Baillière et Fils |volume=II |page=Plate VIII |url=https://gallica.bnf.fr/ark:/12148/bpt6k6538959f/f171.item |access-date=13 August 2020 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021313/https://gallica.bnf.fr/ark:/12148/bpt6k6538959f/f171.item |url-status=live }}</ref>
*''Panthera gombaszoegensis'' (])
*''P. o. hernandesii'' {{small|(], 1857)}} was a jaguar from ] in Mexico.<ref>{{cite journal |author=Gray, J. E. |year=1857 |title=Notice of a new species of jaguar from Mazatlan, living in the gardens of the Zoological Society |journal=Proceedings of the Zoological Society of London |volume=25 |page=278 |url=https://archive.org/details/lietuvostsrmoksl56liet/page/278/mode/2up}}</ref>
*''P. o. palustris'' {{small|(], 1888)}} was a ] jaguar ] excavated in the ] of ], Argentina.<ref>{{cite book |author=Ameghino, F. |year=1888 |chapter=Formación Pampeana |title=Los Mamíferos fósiles de la República Argentina |language=es |location=Buenos Aires |publisher=Pablo E. Coni é hijos |chapter-url=https://archive.org/details/mamferosfsil00ameg/page/476/mode/1up |pages=473–493}}</ref>
*''P. o. centralis'' {{small|(], 1901)}} was a skull of a male jaguar from ], Costa Rica.<ref name=Mearns>{{cite journal |author=Mearns, E. A. |year=1901 |title=The American Jaguars |journal=Proceedings of the Biological Society of Washington |volume=14 |pages=137–143 |url=https://archive.org/details/3908800952802714biolrich/page/138/mode/2up}}</ref>
*''P. o. goldmani'' {{small|(Mearns, 1901)}} was a jaguar skin from Yohatlan in ], Mexico.<ref name=Mearns />
*''P. o. paraguensis'' {{small|(], 1914)}} was a skull of a male jaguar from Paraguay.<ref>{{cite journal |author=Hollister, N. |year=1915 |title=Two new South American jaguars |journal=Proceedings of the United States National Museum |volume=48 |issue=2069 |pages=169–170 |url=https://archive.org/details/proceedingsofuni481915unit/page/168/mode/2up |doi=10.5479/si.00963801.48-2069.169}}</ref>
*''P. o. arizonensis'' {{small|(], 1932)}} was a skin and skull of a male jaguar from the vicinity of ].<ref name=Goldman1932>{{cite journal |author=Goldman, E. A. |year=1932 |title=The jaguars of North America |journal=Proceedings of the Biological Society of Washington |volume=45 |pages=143–146 |url=https://archive.org/details/proceedingsofbi451932biol/page/142/mode/2up}}</ref>
*''P. o. veraecrucis'' {{small|(] and Goldman, 1933)}} was a skull of a male jaguar from ] in Mexico.<ref>{{cite journal |author1=Nelson, E. W. |author2=Goldman, E. A. |year=1933 |name-list-style=amp |title=Revision of the jaguars |journal=Journal of Mammalogy |volume=14 |issue=3 |pages=221–240 |doi=10.2307/1373821 |jstor=1373821}}</ref>


Reginald Innes Pocock placed the jaguar in the ] ''Panthera'' and observed that it shares several morphological features with the ] (''P. pardus''). He, therefore, concluded that they are most closely related to each other.<ref name=Pocock1939 /> Results of ] and ] research indicate a ] north–south variation between populations, but no evidence for subspecific differentiation.<ref name=Eizirik /><ref name=Larson>{{cite journal |last=Larson |first=S. E. |year=1997 |title=Taxonomic re-evaluation of the jaguar |journal=] |volume=16 |issue=2 |pages=107–120 |doi=10.1002/(SICI)1098-2361(1997)16:2<107::AID-ZOO2>3.0.CO;2-E}}</ref> ] analysis of 84 jaguar samples from South America revealed that the ] between jaguar populations in ] was high in the past.<ref>{{cite journal |author1=Ruiz-Garcia, M. |author2=Payan, E. |author3=Murillo, A. |author4=Alvarez, D. |year=2006 |name-list-style=amp |title=DNA microsatellite characterization of the jaguar (''Panthera onca'') in Colombia |journal=Genes & Genetic Systems |volume=81 |issue=2 |pages=115–127 |url=https://www.jstage.jst.go.jp/article/ggs/81/2/81_2_115/_pdf |doi=10.1266/ggs.81.115 |doi-access=free |pmid=16755135 |access-date=13 December 2014 |archive-date=16 December 2014 |archive-url=https://web.archive.org/web/20141216093206/https://www.jstage.jst.go.jp/article/ggs/81/2/81_2_115/_pdf |url-status=live }}</ref> Since 2017, the jaguar is considered to be a ],<ref name="catsg">{{cite journal |author1=Kitchener, A. C. |author2=Breitenmoser-Würsten, C. |author3=Eizirik, E. |author4=Gentry, A. |author5=Werdelin, L. |author6=Wilting, A. |author7=Yamaguchi, N. |author8=Abramov, A. V. |author9=Christiansen, P. |author10=Driscoll, C. |author11=Duckworth, J. W. |author12=Johnson, W. |author13=Luo, S.-J. |author14=Meijaard, E. |author15=O'Donoghue, P. |author16=Sanderson, J. |author17=Seymour, K. |author18=Bruford, M. |author19=Groves, C. |author20=Hoffmann, M. |author21=Nowell, K. |author22=Timmons, Z. |author23=Tobe, S. |year=2017 |name-list-style=amp |title=A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group |journal=Cat News |volume=Special Issue 11 |pages=70–71 |url=https://repository.si.edu/bitstream/handle/10088/32616/A_revised_Felidae_Taxonomy_CatNews.pdf?sequence=1&isAllowed=y#page=70 |access-date=13 May 2018 |archive-date=30 July 2018 |archive-url=https://web.archive.org/web/20180730142355/https://repository.si.edu/bitstream/handle/10088/32616/A_revised_Felidae_Taxonomy_CatNews.pdf?sequence=1&isAllowed=y#page=70 |url-status=live }}</ref> though the modern ''Panthera onca onca'' is still distinguished from two fossil subspecies, ''Panthera onca augusta'' and ''Panthera onca mesembrina''. However, the 2024 study suggested that the validity of subspecific assignments on both ''P. o. augusta'' and ''P. o. mesembrina'' remains unresolved, since both fossil and living jaguars show a considerable variation in ].<ref>{{cite journal|author1=Srigyan, M.|author2=Schubert, B.W.|author3=Bushell, M.|author4=Santos, S.H.D.|author5=Figueiró, H.V.|author6=Sacco, S.|author7=Eizirik, E.|author8=Shapiro, B.|year=2024|title=Mitogenomic analysis of a late Pleistocene jaguar from North America|journal=Journal of Heredity|volume=115|issue=4|pages=424–431|doi=10.1093/jhered/esad082|pmid=38150503 |pmc=11235123}}</ref>
==Ecological role==
]]]The ecological role of the jaguar most closely resembles the tiger. They are considered an umbrella species. An umbrella species is defined as a species that generally covers large areas in their daily or seasonal movements. They serve as "mobile links" at the landscape scale, through predation, seed dispersal or pollination. Protecting enough habitat and connectivity to assure viable population of these organisms benefits many other species more restricted in their range. The jaguar is an ]. They can run at amazingly high speeds, using its speed to catch its prey.


==Prey== ===Evolution===
]
<!-- Image with unknown copyright status removed: ] -->
The ''Panthera'' ] is estimated to have ] from the ] of the ] around {{Ma|9.32|4.47|million years ago}} to {{Ma|11.75|0.97|million years ago}}.<ref name="Johnson2006">{{cite journal |last1=Johnson |first1=W. E. |last2=Eizirik |first2=E. |last3=Pecon-Slattery |first3=J. |last4=Murphy |first4=W. J. |last5=Antunes |first5=A. |last6=Teeling |first6=E. |last7=O'Brien |first7=S. J. |name-list-style=amp |title=The late miocene radiation of modern Felidae: A genetic assessment |journal=] |volume=311 |issue=5757 |pages=73–77 |date=2006 |pmid=16400146 |doi=10.1126/science.1122277 |bibcode=2006Sci...311...73J |s2cid=41672825 |url=https://zenodo.org/record/1230866 |access-date=24 November 2021 |archive-date=4 October 2020 |archive-url=https://web.archive.org/web/20201004075725/https://zenodo.org/record/1230866 |url-status=live }}</ref><ref name="Werdelin2010">{{cite book |last1=Werdelin |first1=L. |last2=Yamaguchi |first2=N. |last3=Johnson |first3=W. E. |last4=O'Brien |first4=S. J. |name-list-style=amp |chapter=Phylogeny and evolution of cats (Felidae) |date=2010 |pages=59–82 |publisher=Oxford University Press |location=Oxford, UK |isbn=978-0-19-923445-5 |chapter-url=https://www.researchgate.net/publication/266755142 |editor1-last=Macdonald |editor1-first=D. W. |editor2-last=Loveridge |editor2-first=A. J. |title=Biology and Conservation of Wild Felids |access-date=24 November 2021 |archive-date=25 September 2018 |archive-url=https://web.archive.org/web/20180925141956/https://www.researchgate.net/publication/266755142 |url-status=live }}</ref><ref name=Li_al2016>{{cite journal |last1=Li |first1=G. |last2=Davis |first2=B. W. |last3=Eizirik |first3=E. |last4=Murphy |first4=W. J. |name-list-style=amp |date=2016 |title=Phylogenomic evidence for ancient hybridization in the genomes of living cats (Felidae) |journal=Genome Research |volume=26 |issue=1 |pages=1–11 |doi=10.1101/gr.186668.114 |pmid=26518481 |pmc=4691742}}</ref> Some genetic analyzes place the jaguar as a ] to the lion with which it diverged {{Ma|3.46|1.22|million years ago}},<ref name=Johnson2006/><ref name="Werdelin2010"/> but other studies place the lion closer to the leopard.<ref name="davis2010">{{cite journal |last1=Davis |first1=B. W. |last2=Li |first2=G. |last3=Murphy |first3=W. J. |name-list-style=amp |title=Supermatrix and species tree methods resolve phylogenetic relationships within the big cats, ''Panthera'' (Carnivora: Felidae) |journal=Molecular Phylogenetics and Evolution |year=2010 |volume=56 |issue=1 |pages=64–76 |doi=10.1016/j.ympev.2010.01.036 |pmid=20138224 |bibcode=2010MolPE..56...64D |url=https://www.academia.edu/12157986 |access-date=24 November 2021 |archive-date=21 November 2021 |archive-url=https://web.archive.org/web/20211121100453/https://www.academia.edu/12157986 |url-status=live }}</ref><ref name="mazak2011">{{cite journal |last1=Mazák|first1=J. H. |last2=Christiansen |first2=P. |last3=Kitchener |first3=A. C. |last4=Goswami |first4=A. |name-list-style=amp |title=Oldest known pantherine skull and evolution of the tiger |journal=PLOS ONE |year=2011 |volume=6 |issue=10 |pages=e25483 |doi=10.1371/journal.pone.0025483 |pmid=22016768 |pmc=3189913 |bibcode=2011PLoSO...625483M|doi-access=free}}</ref>
Jaguars are hunters that do not work with one another outside the breeding season. They hunt around 85 different species including: ], ]s, ], and even ], up to a certain size. They are opportunists and will take anything from frogs, mice, birds, and fish to domestic livestock. A jaguar's bite can pierce the shell of a ] (Emmons, 1987). Jaguars are considered a stalk and ambush predator and are not meant to run over long distances but prefer to surprise unsuspecting prey.


The lineage of the jaguar appears to have originated in Africa and spread to Eurasia 1.95–1.77 mya. The living jaguar species is often suggested to have descended from the Eurasian ''].'' The ancestor of the jaguar entered the American continent via ], the land bridge that once spanned the ],<ref name="Argant2011">{{cite journal |first1=A. |last1=Argant |first2=J. |last2=Argant |name-list-style=amp |title=The ''Panthera gombaszogensis'' story: the contribution of the Château Breccia (Saône-et-Loire, Burgundy, France) |journal=Quaternaire |issue=Hors-serie 4 |year=2011 |pages=247–269 |url=https://www.researchgate.net/publication/286036249 |access-date=24 November 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021307/https://www.researchgate.net/publication/286036249_The_Panthera_Gombaszogensis_story_The_contribution_of_the_chateau_breccia_Saone-Et-Loire_Burgundy_France |url-status=live }}</ref><ref>{{cite journal |last1=Jiangzuo|first1=Q. |last2=Liu |first2=J. |name-list-style=amp |year=2020|title=First record of the Eurasian jaguar in southern Asia and a review of dental differences between pantherine cats|journal=Journal of Quaternary Science |volume=35|issue=6|pages=817–830 |doi=10.1002/jqs.3222|bibcode=2020JQS....35..817J |s2cid=219914902}}</ref> Some authors have disputed the close relationship between ''P. gombaszoegensis'' (which is primarily known from Europe) and the modern jaguar.<ref name=":1">{{Cite journal |last1=Chatar |first1=Narimane |last2=Michaud |first2=Margot |last3=Fischer |first3=Valentin |date=September 2022 |editor-last=Silcox |editor-first=Mary |title=Not a jaguar after all? Phylogenetic affinities and morphology of the Pleistocene felid Panthera gombaszoegensis |url=https://onlinelibrary.wiley.com/doi/10.1002/spp2.1464 |journal=Papers in Palaeontology |language=en |volume=8 |issue=5 |doi=10.1002/spp2.1464 |bibcode=2022PPal....8E1464C |hdl=2268/294237 |issn=2056-2799 |s2cid=252489047 |hdl-access=free |access-date=13 March 2024 |archive-date=27 September 2022 |archive-url=https://web.archive.org/web/20220927093151/https://onlinelibrary.wiley.com/doi/10.1002/spp2.1464 |url-status=live }}</ref> The oldest fossils of modern jaguars (''P. onca'') have been found in North America dating between 850,000-820,000 years ago.<ref name=SK1993/> Results of ] analysis of 37 jaguars indicate that current populations evolved between 510,000 and 280,000 years ago in northern South America and subsequently recolonized North and Central America after the extinction of jaguars there during the ].<ref name=Eizirik>{{cite journal |author1=Eizirik, E. |author2=Kim, J. H. |author3=Menotti-Raymond, M. |author4=Crawshaw P. G. Jr. |author5=O'Brien, S. J. |author6=Johnson, W. E. |year=2001 |name-list-style=amp |title=Phylogeography, population history and conservation genetics of jaguars (''Panthera onca'', Mammalia, Felidae) |journal=Molecular Ecology |volume=10 |issue=1 |pages=65–79 |doi=10.1046/j.1365-294X.2001.01144.x |pmid=11251788 |bibcode=2001MolEc..10...65E |s2cid=3916428 |url=https://zenodo.org/record/1236534 }}</ref>
The jaguar uses a different killing method than most cats to kill its prey. Instead of biting the neck, to suffocate or sever the spinal cord, the jaguar delivers a fatal bite directly to the ], piercing the brain. It is because of this killing technique that jaguars often break teeth as they progress in age. Jaguars eat from 10-70 pounds (5-32 kg) of food daily.


Two extinct subspecies of jaguar are recognized in the fossil record: the North American '']'' and South American '']''.<ref>{{cite journal|last1=Chahud|first1=A. |last2=Okumura|first2=M. |year=2020|title=The presence of ''Panthera onca'' Linnaeus 1758 (Felidae) in the Pleistocene of the region of Lagoa Santa, State of Minas Gerais, Brazil |journal=Historical Biology |volume=33|issue=10|pages=2496–2503 |doi=10.1080/08912963.2020.1808975 |s2cid=225408043}}</ref>
==Predators==
{{clade gallery |align=left |width=520px; |main-caption=] relationships of the jaguar as derived through analysis of
Jaguars have very few predators. Their main predators are humans, but attacks by large anacondas and crocodilians have been documented.
|header1=nuclear DNA:<ref name=Johnson2006 />
|cladogram1={{clade |label1=Felidae
|1={{clade |label2=Pantherinae
|1=Felinae ]
|2={{clade |label1=''Panthera''
|1={{clade
|1={{clade
|1={{clade
|1=] (''P. leo'') ]
|2='''Jaguar''' ]}}
|2=] (''P. pardus'') ]}}
|2={{clade
|1=] (''P. tigris'') ]
|2=] (''P. uncia'') ]}} }}
|2='']'' ]}} }} }}
|header2=mitochondrial DNA:<ref name=Li_al2016 />
|cladogram2={{clade |label1=Felidae
|1={{clade |label2=Pantherinae
|1=Felinae ]
|2={{clade |label1=''Panthera''
|1={{clade
|1={{clade
|1={{clade
|1={{clade|1=Lion ]|2=Leopard ]}}
|2=Snow leopard ]}}
|2='''Jaguar''' ]}}
|2=Tiger ]}}
|2=''Neofelis'' ]}} }} }}
}}{{clear|left}}


==Reproduction== ==Description==
{{multiple image |direction=vertical |align=right
]
|image1=Description iconographique comparée du squelette et du système dentaire des mammifères récents et fossiles (Panthera onca skull).jpg |caption1=Skull of a jaguar
Young jaguar males reach sexual maturity at about three to four years of age, with females reaching maturity about a year earlier than males. Females give birth to as many as four cubs after a 90 to 110 day gestation, but raise no more than two of them to adulthood. The young are born blind and can see after two weeks. They remain with their mother for up to two years before leaving to establish a territory for themselves, which can be anywhere between 25 and 150 square kilometers in size (depending on the availability of suitable prey). Typical lifespan is 10 or 11 years in the wild; in captivity, jaguars have lived up to 20 years.
|image2=Cheetah, leopard & jaguar (en).jpg |caption2=Illustration of ], leopard and jaguar
|image3=Black Jaguar (Panthera onca).JPG |caption3=A black jaguar. Such ] jaguars as well as leopards are commonly called ]s}}


The jaguar is a compact and muscular animal. It is the largest cat native to the Americas and the ] in the world, exceeded in size only by the tiger and the lion.<ref name=Seymour /><ref name="sizes">{{cite journal |last1=Hayward |first1=M. W. |last2=Kamler |first2=J. F. |last3=Montgomery |first3=R. A. |last4=Newlove |first4=A. |year=2016 |name-list-style=amp |title=Prey Preferences of the Jaguar ''Panthera onca'' Reflect the Post-Pleistocene Demise of Large Prey |journal=Frontiers in Ecology and Evolution |volume=3 |page=148 |doi=10.3389/fevo.2015.00148 |doi-access=free}}</ref><ref name="sizes22">{{cite journal |last1=Hope |first1=M. K. |last2=Deem |first2=S. L. |year=2006 |name-list-style=amp |title=Retrospective Study of Morbidity and Mortality of Captive Jaguars (''Panthera onca'') in North America: 1982–2002 |journal=Zoo Biology |volume=25 |issue=6 |pages=501–512 |url=https://repository.si.edu/bitstream/handle/10088/11687/Zoo%20Biology%2C%20Vol.%2025%2C%20Issue%206%20Retrospective%20Study%20of%20Morbidity%20and%20Mortality%20of%20Captive%20Jaguars....pdf?sequence=1&isAllowed=y |doi=10.1002/zoo.20112 |access-date=12 September 2018 |archive-date=9 September 2018 |archive-url=https://web.archive.org/web/20180909000241/https://repository.si.edu/bitstream/handle/10088/11687/Zoo%20Biology%2C%20Vol.%2025%2C%20Issue%206%20Retrospective%20Study%20of%20Morbidity%20and%20Mortality%20of%20Captive%20Jaguars....pdf?sequence=1&isAllowed=y |url-status=live }}</ref> It stands {{cvt|57|to|81|cm|1}} tall at the shoulders.<ref>{{cite magazine |author1=Rich, M.S. |year=1976 |title=The jaguar |magazine=Zoonoz |volume=49 |issue=9 |pages=14–17}}</ref><ref name=Scognamillo2003>{{cite journal |author1=Scognamillo, D. |author2=Maxit, I. E. |author3=Sunquist, M. |author4=Polisar, J. |year=2003 |name-list-style=amp |title=Coexistence of jaguar (''Panthera onca'') and puma (''Puma concolor'') in a mosaic landscape in the Venezuelan llanos |journal=Journal of Zoology |volume=259 |issue=3 |pages=269–279 |doi=10.1017/S0952836902003230}}</ref>
==The jaguar in Central and South American culture==
Its size and weight vary considerably depending on sex and region: weights in most regions are normally in the range of {{cvt|56|-|96|kg}}. Exceptionally big males have been recorded to weigh as much as {{cvt|158|kg}}.<ref name=r10 /><ref name="Animal">{{cite book |author1=Burnie, D. |author2=Wilson, D.E. |year=2001 |name-list-style=amp |title=Animal: The Definitive Visual Guide to the World's Wildlife |location=New York City |publisher=] |isbn=978-0-7894-7764-4}}</ref>
:''See also: ]''
The smallest females from ] weigh about {{cvt|36|kg}}. It is ], with females typically being 10–20% smaller than males. The length from the nose to the base of the tail varies from {{cvt|1.12|to|1.85|m|ftin}}. The tail is {{cvt|45|to|75|cm}} long and the shortest of any ].<ref name=r10>{{cite book |author=Nowak, R.M. |year=1999 |title=Walker's Mammals of the World |publisher=] |location=Baltimore |volume=2 |page=831 |url={{Google books|T37sFCl43E8C|page=PA831|keywords=|text=|plainurl=yes}} |isbn=978-0-8018-5789-8}}</ref>
The word ''jaguar'' comes from the ] ] language. According to one early ] explorer, ''jaguara'' meant "a beast that kills its prey with one bound." However, this is incorrect. ''Jaguara'' refers to any carnivorous animal. The original and complete indigenous name for the species is ''Yaguareté'', where -''eté'' means "true". Curiously, ''Yagua'' means "fierce" in ].
Its muscular legs are shorter than the legs of other ''Panthera'' species with similar body weight.<ref name=Gonyea1976 />
Jaguar is also a royal title bestowed to a royal prince, princess or ruling monarch in some ] traditions such as that of the ].


Size tends to increase from north to south. Jaguars in the ] on the Pacific coast of central Mexico weighed around {{cvt|50|kg}}.<ref name=foodhabits>{{cite journal |author1=Nuanaez, R. |author2=Miller, B. |author3=Lindzey, F. |year=2000 |name-list-style=amp |title=Food habits of jaguars and pumas in Jalisco, Mexico |journal=] |volume=252 |issue=3 |pages=373–379 |url=http://journals.cambridge.org/action/displayAbstract?fromPage=online&aid=58851 |doi=10.1111/j.1469-7998.2000.tb00632.x |access-date=8 September 2006 |archive-date=10 August 2016 |archive-url=https://web.archive.org/web/20160810211518/http://journals.cambridge.org/action/displayAbstract?fromPage=online&aid=58851 |url-status=live }}</ref>
In Central and South America the jaguar has long been a symbol of power and strength. To the ] ], the jaguar was believed to communicate between the living and the dead, as well as protect the royal household. The Maya saw these powerful felines as their companions in the spiritual world. The Aztec civilization also had the same image of the jaguar as the representative of the ruler and as a warrior. The ]s formed an elite warrior class known as the jaguar knights. In ], the jaguar was also considered to be the ] animal of the powerful deity ].
Jaguars in Venezuela and ] are much larger, with average weights of about {{cvt|95|kg}} in males and of about {{cvt|56|-|78|kg}} in females.<ref name=Seymour />


The jaguar's ] ranges from pale yellow to tan or reddish-yellow, with a whitish underside and covered in black spots. The spots and their shapes vary: on the sides, they become ] which may include one or several dots. The spots on the head and neck are generally solid, as are those on the tail where they may merge to form bands near the end and create a black tip. They are elongated on the middle of the back, often connecting to create a median stripe, and blotchy on the belly.<ref name=Seymour /> These patterns serve as ] in areas with dense vegetation and patchy shadows.<ref>{{cite journal |author1=Allen, W.L. |author2=Cuthill, I.C. |author3=Scott-Samuel, N.E. |author4=Baddeley, R. |year=2010 |name-list-style=amp |title=Why the leopard got its spots: relating pattern development to ecology in felids |journal=Proceedings of the Royal Society B |volume=278 |issue=1710 |pages=1373–1380 |doi=10.1098/rspb.2010.1734 |pmc=3061134 |pmid=20961899}}</ref>
==Melanism==
Jaguars living in forests are often darker and considerably smaller than those living in open areas, possibly due to the smaller numbers of large, herbivorous prey in forest areas.<ref name=CAP>{{cite book |author1=Nowell, K. |author2=Jackson, P. |year=1996 |name-list-style=amp |chapter=Jaguar, ''Panthera onca'' (Linnaeus, 1758) |title=Wild Cats. Status Survey and Conservation Action Plan |publisher=IUCN/SSC Cat Specialist Group |location=Gland, Switzerland |chapter-url=http://carnivoractionplans1.free.fr/wildcats.pdf#page=143 |pages=118–122 |access-date=31 August 2006 |archive-date=7 August 2007 |archive-url=https://web.archive.org/web/20070807215533/http://carnivoractionplans1.free.fr/wildcats.pdf#page=143 |url-status=live }}</ref>
]
The background of the coat is usually an orange-yellow in colour, with numerous rings or rosettes on the flanks and spots on the head and neck. A condition known as ] occasionally occurs and can create jaguars that appear entirely black (although the spots are still visible if one looks closely). These are known as ]s, but do not form a separate species.


The jaguar closely resembles the leopard but is generally more robust, with stockier limbs and a more square head. The rosettes on a jaguar's coat are larger, darker, fewer in number and have thicker lines, with a small spot in the middle.<ref name=Gonyea1976>{{cite journal |last=Gonyea |first=W.J. |year=1976 |title=Adaptive differences in the body proportions of large felids |journal=Acta Anatomica |volume=96 |issue=1 |pages=81–96 |doi=10.1159/000144663 |pmid=973541}}</ref>
==Hybridization==
It has powerful jaws with the third-highest bite force of all felids, after the tiger and the lion.<ref name=Wroe2006>{{cite journal |author1=Wroe, S. |author2=McHenry, C. |author3=Thomason, J. |year=2006 |name-list-style=amp |title=Bite club: comparative bite force in big biting mammals and the prediction of predatory behavior in fossil taxa |journal=Proceedings of the Royal Society B |volume=272 |issue=1563 |pages=619–625 |doi=10.1098/rspb.2004.2986 |doi-access=free |pmc=1564077 |pmid=15817436}}</ref>
Jaguars are occasionally mated with other big cats such as the lion, tiger and leopard. These hybridizations are usually carried out in controlled environments. For more information on hybrid cats see ].
It has an average bite force at the ] tip of 887.0 ] and a ] at the canine tip of 118.6.<ref>{{cite journal |author=Christiansen, P. |year=2007 |title=Canine morphology in the larger Felidae: implications for feeding ecology |journal=Biological Journal of the Linnean Society |volume=91 |issue=4 |pages=573–592 |doi=10.1111/j.1095-8312.2007.00819.x |doi-access=free}}</ref>
A {{cvt|100|kg}} jaguar can bite with a force of {{cvt|4.939|kN|lbf}} with the canine teeth and {{cvt|6.922|kN|lbf}} at the ] notch.<ref>{{cite journal |last1=Hartstone-Rose |first1=A. |last2=Perry |first2=J.M.G. |last3=Morrow |first3=C.J. |year=2012 |name-list-style=amp |title=Bite Force Estimation and the Fiber Architecture of Felid Masticatory Muscles |journal=The Anatomical Record: Advances in Integrative Anatomy and Evolutionary Biology |volume=295 |issue=8 |pages=1336–1351 |doi=10.1002/ar.22518 |doi-access= |pmid=22707481 |s2cid=35304260}}</ref>


===Color variation===
==Trivia==
] jaguars are also known as ]s. The black ] is less common than the spotted one.<ref>{{cite book |author1=Brown, D.E. |author2=Lopez-Gonzalez, C.A. |date=2001 |name-list-style=amp |title=Borderland jaguars: tigres de la frontera |publisher=University of Utah Press |location=Salt Lake City, UT}}</ref>
The crushing force of a jaguar's jaws is 600 psi (41 bar), according to Discovery channel. However, little is known about the method of measurement. The result of this measurement was used to construct simulated species for the ] series '']''.
Black jaguars have been documented in Central and South America. Melanism in the jaguar is caused by deletions in the ] gene and inherited through a ].<ref>{{cite journal |author1=Eizirik, E. |author2=Yuhki, N. |author3=Johnson, W.E. |author4=Menotti-Raymond, M. |author5=Hannah, S.S. |author6=O'Brien, S.J. |year=2003 |name-list-style=amp |title=Molecular Genetics and Evolution of Melanism in the Cat Family |journal=Current Biology |volume=13 |issue=5 |pages=448–453 |doi=10.1016/S0960-9822(03)00128-3 |pmid=12620197 |s2cid=19021807 |doi-access=free|bibcode=2003CBio...13..448E }}</ref> Black jaguars occur at higher densities in tropical rainforest and are more active during the daytime. This suggests that melanism provides camouflage in dense vegetation with high illumination.<ref name=Mooring2020/>


In 2004, a camera trap in the ] mountains photographed the first documented black jaguar in Northern Mexico.<ref name=Dinets>{{cite journal |last1=Dinets |first1=V. |last2=Polechla |first2=P.J. |year=2005 |name-list-style=amp |title=First documentation of melanism in the jaguar (''Panthera onca'') from northern Mexico |journal=Cat News |volume=42 |page=18 |url=http://dinets.travel.ru/blackjaguar.htm |url-status=dead |archive-url=https://web.archive.org/web/20060926024755/http://dinets.travel.ru/blackjaguar.htm |archive-date=26 September 2006}}</ref> Black jaguars were also photographed in Costa Rica's ], in the mountains of the ], in ] and in eastern ].<ref>{{cite journal |author1=Núñez, M.C. |author2=Jiménez, E.C. |year=2009 |name-list-style=amp |title=A new record of a black jaguar, ''Panthera onca'' (Carnivora: Felidae) in Costa Rica |journal=Brenesia |volume=71 |pages=67–68 |url=https://www.researchgate.net/publication/313473228 |access-date=12 April 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021258/https://www.researchgate.net/publication/313473228_New_record_of_a_black_jaguar_Panthera_onca_Carnivora_Felidae_in_Costa_Rica |url-status=live }}</ref><ref name=Mooring2020>{{cite journal |author1=Mooring, M. S. |author2=Eppert, A. A. |author3=Botts, R. T. |year=2020 |name-list-style=amp |title=Natural Selection of Melanism in Costa Rican Jaguar and Oncilla: A Test of Gloger's Rule and the Temporal Segregation Hypothesis |journal=Tropical Conservation Science |volume=13 |pages=1–15 |doi=10.1177/1940082920910364 |doi-access=free}}</ref><ref>{{cite journal |author1=Sáenz-Bolaños, C. |name-list-style=amp |author2=Montalvo, V. |author3=Fuller, T.K. |author4=Carrillo, E. |year=2015 |title=Records of black jaguars at Parque Nacional Barbilla, Costa Rica |journal=Cat News |issue=62 |pages=38–39}}</ref><ref>{{cite journal |author1=Yacelga, M. |name-list-style=amp |author2=Craighead, K. |year=2019 |title=Melanistic jaguars in Panama |journal=Cat News |issue=70 |pages=39–41 |url=https://www.academia.edu/41977728 |access-date=17 April 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021257/https://www.academia.edu/41977728 |url-status=live }}</ref>
==Jaguars in Culture==
The name ] is given to the brand of a luxury ].


==Distribution and habitat==
The jaguar is the mascot of the ]'s ] franchise.
{{multiple image |direction=vertical |align=right
|image1=Jaguar (Panthera onca palustris) female Piquiri River.JPG |caption1=A female jaguar at ], ] state, Brazil
|image2=Jaguar (Panthera onca) male back in the water (29173428825).jpg |caption2=A jaguar in ]}}


In 1999, the jaguar's historic range at the turn of the 20th century was estimated at {{cvt|19000000|km2}}, stretching from the southern United States through Central America to southern Argentina. By the turn of the 21st century, its global range had decreased to about {{cvt|8750000|km2}}, with most declines occurring in the southern United States, northern Mexico, northern Brazil, and southern Argentina.<ref name=Sanderson2002>{{cite journal |last1=Sanderson |first1=E. W. |last2=Redford |first2=K. H. |last3=Chetkiewicz |first3=C. L. B. |last4=Medellin |first4=R. A. |last5=Rabinowitz |first5=A. R. |author5-link=Alan Rabinowitz |last6=Robinson |first6=J. G. |last7=Taber |first7=A. B. |year=2002 |name-list-style=amp |title=Planning to save a species: the jaguar as a model |journal=Conservation Biology |volume=16 |issue=1 |pages=58–72 |doi=10.1046/j.1523-1739.2002.00352.x |pmid=35701976 |s2cid=3955250 |doi-access=free |bibcode=2002ConBi..16...58S }}</ref>
Atari's last video game console was the ].
Its present range extends from Mexico through Central America to South America comprising ], ], ], ], ], particularly on the ], ], ], ], ], ], ], ], ], ], Brazil, ] and ]. It is considered to be ] in ] and ].<ref name=iucn />


Jaguars have been occasionally sighted in ], ] and ], with 62 accounts reported in the 20th century.<ref name=BG2000>{{cite journal |author1=Brown, D. E. |author2=González, C. A. L. |year=2000 |name-list-style=amp |title=Notes on the occurrences of jaguars in Arizona and New Mexico |journal=The Southwestern Naturalist |volume=45 |issue=4 |pages=537–542 |doi=10.2307/3672607 |jstor=3672607}}</ref><ref name=Pavlik>{{Cite journal |last=Pavlik |first=S. |year=2003 |title=Rohonas and spotted Lions: The historical and cultural occurrence of the Jaguar, ''Panthera onca'', among the native tribes of the American Southwest |journal=Wíčazo Ša Review |volume=18 |issue=1 |pages=157–175 |doi=10.1353/wic.2003.0006 |jstor=1409436 |s2cid=161236104}}</ref>
Mac OS X 10.2 had the codename "Jaguar."
Between 2012 and 2015, a male ] jaguar was recorded in 23 locations in the ].<ref>{{cite report |author=Culver, M. |title=Open-File Report |year=2016 |chapter=Jaguar surveying and monitoring in the United States |publisher=U.S. Geological Survey |location=Reston, VA |series=2016-1095 |doi=10.3133/ofr20161095 |doi-access=free}}</ref> Eight jaguars were photographed in the southwestern US between 1996 and 2024.<ref>{{cite web|date=2024|title=Endangered jaguar previously unknown to U.S. is caught on camera in Arizona|website=CBS News|url=https://www.cbsnews.com/news/endangered-jaguar-spotted-in-arizona-previously-unknown-to-us/|access-date=June 21, 2024}}</ref>


The jaguar prefers dense forest and typically inhabits dry ]s, ], ]s and ]s in Central and South America; open, seasonally flooded ]s, dry ] and historically also ]s in the United States. It has been recorded at elevations up to {{cvt|3800|m}} but avoids ]s. It favors riverine habitat and ]s with dense vegetation cover.<ref name=CAP /> In the Mayan forests of Mexico and Guatemala, 11 GPS-collared jaguars preferred undisturbed dense habitat away from roads; females avoided even areas with low levels of human activity, whereas males appeared less disturbed by human population density.<ref>{{cite journal |author1=Colchero, F. |author2=Conde, D. A. |author3=Manterola, C. |author4=Chávez, C. |author5=Rivera, A. |author6=Ceballos, G. |year=2011 |name-list-style=amp |title=Jaguars on the move: modeling movement to mitigate fragmentation from road expansion in the Mayan Forest |journal=Animal Conservation |volume=14 |issue=2 |pages=1–9 |doi=10.1111/J.1469-1795.2010.00406.X |bibcode=2011AnCon..14..158C |s2cid=62820463 |url=https://www.demogr.mpg.de/publications/files/4097_1300970681_1_ArticlePdf.pdf |access-date=14 March 2021 |archive-date=26 January 2021 |archive-url=https://web.archive.org/web/20210126211109/https://www.demogr.mpg.de/publications/files/4097_1300970681_1_ArticlePdf.pdf |url-status=live}}</ref> A young male jaguar was also recorded in the ] ] at a waterhole.<ref>{{cite journal |author1=Caso, A. |name-list-style=amp |author2=Domínguez, E. F. |year=2018 |title=Confirmed presence of jaguar, ocelot and jaguarundi in the Sierra of San Carlos, Mexico |journal=Cat News |issue=68 |pages=31–32}}</ref>
The jaguar is the name of a Mexican soccer national league team: ].


=== Former range ===
The jaguar serves as the mascot for the following colleges & universities: ]-Baton Rouge, ], University of Southern Alabama.
In the 19th century, the jaguar was still sighted at the ] {{cvt|30|-|50|mile|km|order=flip}} north of ] in ], in coastal ], northern Arizona and New Mexico.<ref name=Daggett>{{cite journal |author1=Daggett, P. M. |author2=Henning, D. R. |year=1974 |name-list-style=amp |title=The Jaguar in North America |journal=American Antiquity |volume=39 |issue=3 |pages=465–469 |doi=10.2307/279437 |jstor=279437|s2cid=160927286}}</ref>
Multiple verified zoological reports of the jaguar are known in California, two as far north as ] in 1814 and 1826. The only record of an active jaguar den with breeding adults and kittens in the United States was in the ] of California prior to 1860.<ref>{{Cite journal |author=Merriam, C.H. |author-link=Clinton Hart Merriam |year=1919 |title=Is the Jaguar entitled to a place in the Californian fauna? |journal=Journal of Mammalogy |volume=1 |issue=1 |pages=38–42 |doi=10.1093/jmammal/1.1.38 |url=https://academic.oup.com/jmammal/article-abstract/1/1/38/875846?redirectedFrom=fulltext |access-date=18 November 2018 |archive-date=18 November 2018 |archive-url=https://web.archive.org/web/20181118210048/https://academic.oup.com/jmammal/article-abstract/1/1/38/875846?redirectedFrom=fulltext |url-status=live}}</ref> The jaguar persisted in California until about 1860.<ref name=Pavlik/>
The last confirmed jaguar in Texas was shot in 1948, {{cvt|3|mile|km|order=flip}} southeast of ].<ref>{{Cite book |last1=Schmidly |first1=D. J. |last2=Bradley |first2=R. D. |date=2016 |title=The Mammals of Texas |publisher=University of Texas Press |edition=Seventh |url=https://www.depts.ttu.edu/nsrl/mammals-of-texas-online-edition/Accounts_Extinct_Carnivora/Panthera_onca.php |place=Lubbock |doi=10.7560/308868 |isbn=9781477310021 |access-date=9 October 2023 |archive-date=18 January 2023 |archive-url=https://web.archive.org/web/20230118025353/https://www.depts.ttu.edu/nsrl/mammals-of-texas-online-edition/Accounts_Extinct_Carnivora/Panthera_onca.php |url-status=live }}</ref>
In Arizona, a female was shot in the ] in 1963. By the late 1960s, the jaguar was thought to have been extirpated in the United States. Arizona outlawed jaguar hunting in 1969, but by then no females remained, and over the next 25 years only two males were sighted and killed in the state. In 1996, a rancher and hunting guide from ] came across a jaguar in the ] and became a researcher on jaguars, placing trail cameras, which recorded four more jaguars.<ref>{{cite journal |title=Return of the Jaguar? |author=Rizzo, W. |journal=] |date=2005 |url=http://www.smithsonianmag.com/ist/?next=/science-nature/return-of-the-jaguar-110630052/ |archive-url=https://web.archive.org/web/20141213225247/http://www.smithsonianmag.com/ist/?next=/science-nature/return-of-the-jaguar-110630052/ |url-status=dead |archive-date=13 December 2014 |access-date=23 November 2011}}</ref>


==Behavior and ecology==
==References==
The jaguar is mostly active at night and during ].<ref name=Scognamillo2003/><ref>{{cite journal |author1=Harmsen, B. J. |author2=Foster, R. J. |author3=Silver, S. C. |author4=Ostro, L. E. T. |author5=Doncaster, C. P. |year=2009 |name-list-style=amp |title=Spatial and temporal interactions of sympatric jaguars (''Panthera onca'') and pumas (''Puma concolor'') in a neotropical forest |journal=Journal of Mammalogy |volume=90 |issue=3 |pages=612–620 |doi=10.1644/08-MAMM-A-140R.1 |doi-access=free}}</ref><ref>{{cite journal |author1=Foster, V.C. |author2=Sarmento, P. |author3=Sollmann, R. |author4=Tôrres, N. |author5=Jácomo, A. T. |author6=Negrões, N. |author7=Fonseca, C. |author8=Silveira, L. |year=2013 |name-list-style=amp |title=Jaguar and Puma Activity Patterns and Predator-Prey Interactions in Four Brazilian Biomes |journal=Biotropica |volume=45 |issue=3 |pages=373–379 |url=https://www.researchgate.net/publication/234153943 |doi=10.1111/btp.12021 |bibcode=2013Biotr..45..373F |s2cid=86338173 |access-date=28 February 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021315/https://www.researchgate.net/publication/234153943_Jaguar_and_Puma_Activity_Patterns_and_Predator-Prey_Interactions_in_Four_Brazilian_Biomes |url-status=live}}</ref>
* {{IUCN2006|assessors=Cat Specialist Group|year=2002|id=15953|title=Panthera onca|downloaded=11 May 2006}} Database entry includes justification for why this species is near threatened
However, jaguars living in densely forested regions of the ] and the ] are largely active by day, whereas jaguars in the ] are primarily active by night.<ref name="Astate et al. 2008">{{cite journal |last1=Astete |first1=S.R. |last2=Sollmann |first2=R. |last3=Silveira |first3=L. |year=2008 |name-list-style=amp |title=Comparative ecology of jaguars in Brazil |journal=Cat News |issue=Special Issue 4 |pages=9–14 |citeseerx=10.1.1.528.3603}}</ref>
The activity pattern of the jaguar coincides with the activity of its main prey species.<ref name="Harmsen et al. 2011">{{cite journal |last1=Harmsen |first1=B.J. |last2=Foster |first2=R.J. |last3=Silver |first3=S. C. |last4=Ostro |first4=L.E.T. |last5=Doncaster |first5=C.P. |year=2011 |name-list-style=amp |title=Jaguar and puma activity patterns in relation to their main prey |journal=Mammalian Biology – Zeitschrift für Säugetierkunde |volume=76 |issue=3 |pages=320–324 |doi=10.1016/j.mambio.2010.08.007|bibcode=2011MamBi..76..320H }}</ref> Jaguars are good swimmers and play and hunt in the water, possibly more than tigers. They have been recorded moving between islands and the shore. Jaguars are also good at climbing trees but do so less often than cougars.<ref name=Seymour />

===Ecological role===
]
The adult jaguar is an ], meaning it is at the top of the ] and is not preyed upon in the wild. The jaguar has also been termed a ], as it is assumed that it controls the population levels of prey such as ] and ] mammals and thus maintains the structural integrity of forest systems.<ref name=foodhabits /><ref>{{cite journal |author=Nijhawan, S. |year=2012 |title=Conservation units, priority areas and dispersal corridors for jaguars in Brazil |journal=Cat News |issue=Special Issue |pages=43–47 |url=http://www.catsg.org/fileadmin/filesharing/5.Cat_News/5.3._Special_Issues/5.3.7._SI_7/Nijhawan_2012_Conservation_units_and_corrdors_for_jaguars_in_Brazil.pdf |access-date=17 August 2020 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021300/http://www.catsg.org/fileadmin/filesharing/5.Cat_News/5.3._Special_Issues/5.3.7._SI_7/Nijhawan_2012_Conservation_units_and_corrdors_for_jaguars_in_Brazil.pdf |url-status=live}}</ref><ref>{{cite journal |author1=Petracca, L.S. |author2=Ramírez-Bravo, O.E. |author3=Hernández-Santín, L. |year=2014 |name-list-style=amp |title=Occupancy estimation of jaguar ''Panthera onca'' to assess the value of east-central Mexico as a jaguar corridor |journal=Oryx |volume=48 |issue=1 |pages=133–140 |doi=10.1017/S0030605313000069 |s2cid=86460403 |doi-access=free}}</ref>
However, field work has shown this may be natural variability, and the population increases may not be sustained. Thus, the ] hypothesis is not accepted by all scientists.<ref name=r30>{{cite journal |author1=Wright, S. J. |author2=Gompper, M. E. |author3=DeLeon, B. |year=1994 |name-list-style=amp |title=Are large predators keystone species in Neotropical forests? The evidence from Barro Colorado Island |journal=Oikos |volume=71 |issue=2 |pages=279–294 |url=https://www.researchgate.net/publication/270356084 |doi=10.2307/3546277 |jstor=3546277 |bibcode=1994Oikos..71..279W |access-date=18 June 2019 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021300/https://www.researchgate.net/publication/270356084_Are_Large_Predators_Keystone_Species_in_Neotropical_Forests_The_Evidence_from_Barro_Colorado_Island |url-status=live}}</ref>

The jaguar is ] with the ]. In central Mexico, both prey on ], which makes up 54% and 66% of jaguar and cougar's prey, respectively.<ref name=foodhabits /> In northern Mexico, the jaguar and the cougar share the same habitat, and their diet overlaps dependent on prey availability. Jaguars seemed to prefer ] and calves. In Mexico and Central America, neither of the two cats are considered to be the dominant predator.<ref name=GGLG2017>{{cite journal |last1=Gutiérrez-González |first1=C. E. |last2=López-González |first2=C. A. |year=2017 |name-list-style=amp |title=Jaguar interactions with pumas and prey at the northern edge of jaguars' range |journal=] |volume=5 |issue=5 |page=e2886 |doi=10.7717/peerj.2886 |pmc=5248577 |pmid=28133569 |doi-access=free}}</ref>
In South America, the jaguar is larger than the cougar and tends to take larger prey, usually over {{cvt|22|kg}}. The cougar's prey usually weighs between {{cvt|2|and|22|kg|0}}, which is thought to be the reason for its smaller size.<ref name=r31>{{cite journal |author1=Iriarte, J. A. |author2=Franklin, W.L. |author3=Johnson, W.E. |author4=Redford, K.H. |year=1990 |name-list-style=amp |title=Biogeographic variation of food habits and body size of the America puma |journal=] |volume=85 |issue=2 |pages=185–190 |bibcode=1990Oecol..85..185I |doi=10.1007/BF00319400 |pmid=28312554 |s2cid=10134066}}</ref>
This situation may be advantageous to the cougar. Its broader prey niche, including its ability to take smaller prey, may give it an advantage over the jaguar in human-altered landscapes.<ref name=foodhabits/>

===Hunting and diet===
]
]]]
The jaguar is an ] and depends solely on flesh for its nutrient requirements. An analysis of 53 studies documenting the diet of the jaguar revealed that its prey ranges in weight from {{cvt|1|to|130|kg}}; it prefers prey weighing {{cvt|45-85|kg}}, with the ] and the ] being the most selected. When available, it also preys on ], ], ] and ].<ref name=sizes/> In floodplains, jaguars opportunistically take reptiles such as ]s and ]s. Consumption of reptiles appears to be more frequent in jaguars than in other big cats.<ref name=reptile>{{cite journal |last1=Miranda |first1=E. |last2=Menezes |first2=J. |last3=Rheingantz |first3=M. L. |year=2016 |name-list-style=amp |title=Reptiles as principal prey? Adaptations for durophagy and prey selection by jaguar (''Panthera onca'') |journal=Journal of Natural History |volume=50 |issue=31–32 |pages=2021–2035 |url=https://zenodo.org/record/3993366 |doi=10.1080/00222933.2016.1180717 |bibcode=2016JNatH..50.2021M |s2cid=89150920 |access-date=28 February 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021309/https://zenodo.org/record/3993366 |url-status=live}}</ref> One remote population in the Brazilian Pantanal is recorded to primarily feed on aquatic reptiles and fish.<ref name=Eriksson2021>{{cite journal|author1=Eriksson, C. |author2=Kantek, D.L. |author3=Miyazaki, S.S. |author4=Morato, R.G. |name-list-style=amp |author5=dos Santos-Filho, M. |author6=Ruprecht, J.S. |author7=Peres, C.A. |author8=Levi, T. |year=2022 |title=Extensive aquatic subsidies lead to territorial breakdown and high density of an apex predator |journal=Ecology |volume=103 |issue=1 |page=e03543 |doi=10.1002/ecy.3543 |doi-access=free |pmid=34841521 |bibcode=2022Ecol..103E3543E |s2cid=242197640}}</ref>
The jaguar also preys on livestock in ] ]ing areas where wild prey is scarce.<ref>{{cite journal |author1=Amit, R. |author2=Gordillo-Chávez, E.J. |author3=Bone, R. |year=2013 |name-list-style=amp |title=Jaguar and puma attacks on livestock in Costa Rica |journal=Human-Wildlife Interactions |volume=7 |issue=1 |pages=77–84}}</ref><ref>{{cite journal |author1=Zarco-González, M.M. |author2=Monroy-Vilchis, O. |author3=Alaníz, J. |year=2013 |name-list-style=amp |title=Spatial model of livestock predation by jaguar and puma in Mexico: conservation planning |journal=Biological Conservation |volume=159 |pages=80–87 |doi=10.1016/j.biocon.2012.11.007|bibcode=2013BCons.159...80Z }}</ref>
The daily food requirement of a captive jaguar weighing {{cvt|34|kg}} was estimated at {{cvt|1.4|kg}} of meat.<ref name=Emmons1987/>

The jaguar's bite force allows it to pierce the ]s of the ] and the ].<ref name=Emmons1987>{{cite journal |author=Emmons, L. H. |year=1987 |title=Comparative feeding ecology of fields in a neotropical rain forest |journal=Behavioral Ecology and Sociobiology |volume=20 |issue=4 |pages=271–283 |url=https://www.researchgate.net/publication/225982805 |doi=10.1007/BF00292180 |bibcode=1987BEcoS..20..271E |s2cid=24990860 |access-date=13 April 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021300/https://www.researchgate.net/publication/225982805_Comparative_Feeding_Ecology_of_Felids_in_a_Neotropical_Rain-Forest |url-status=live}}</ref><ref>{{cite journal |author=Emmons, L. H. |year=1989 |title=Jaguar predation on chelonians |journal=Journal of Herpetology |volume=23 |issue=3 |pages=311–314 |doi=10.2307/1564460 |jstor=1564460}}</ref> It employs an unusual killing method: it bites mammalian prey directly through the ] between the ears to deliver a fatal bite to the brain.<ref name=rosa>{{cite book |last1=Rosa |first1=C. L. de la |last2=Nocke |first2=C. C. |year=2000 |name-list-style=amp |chapter=Jaguar (''Panthera onca'') |title=A guide to the carnivores of Central America: natural history, ecology, and conservation |publisher=] |chapter-url=https://books.google.com/books?id=x5ihAwAAQBAJ&pg=PT39 |pages=39–? |isbn=978-0-292-71604-9 |access-date=28 February 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021333/https://books.google.com/books?id=x5ihAwAAQBAJ&pg=PT39 |url-status=live}}</ref> It kills capybara by piercing its ] through the ]s of its skull, breaking its ] and ] and penetrating its brain, often through the ears.<ref>{{cite journal |author1=Schaller, G.B. |author1-link=George Schaller |author2=Vasconselos, J.M.C. |year=1978 |name-list-style=amp |title=Jaguar predation on capybara |journal=Zeitschrift für Säugetierkunde |volume=43 |pages=296–301 |url=https://www.zobodat.at/pdf/Zeitschrift-Saeugetierkunde_43_0296-0301.pdf |access-date=16 August 2020 |archive-date=25 June 2021 |archive-url=https://web.archive.org/web/20210625121230/https://www.zobodat.at/pdf/Zeitschrift-Saeugetierkunde_43_0296-0301.pdf |url-status=live}}</ref>
It has been hypothesized to be an adaptation to cracking open turtle shells; armored reptiles may have formed an abundant prey base for the jaguar following the late ]s.<ref name=Emmons1987 /> However, this is disputed, as even in areas where jaguars prey on reptiles, they are still taken relatively infrequently compared to mammals in spite of their greater abundance.<ref name=reptile/>

Between October 2001 and April 2004, 10 jaguars were monitored in the southern Pantanal. In the dry season from April to September, they killed prey at intervals ranging from one to seven days; and ranging from one to 16 days in the wet season from October to March.<ref>{{cite journal |author1=Cavalcanti, S. M. C. |author2=Gese, E. M. |year=2010 |name-list-style=amp |title=Kill rates and predation patterns of jaguars (''Panthera onca'') in the southern Pantanal, Brazil |journal=Journal of Mammalogy |volume=91 |issue=3 |pages=722–736 |doi=10.1644/09-MAMM-A-171.1 |doi-access=free}}</ref>

The jaguar uses a stalk-and-ambush strategy when hunting rather than chasing prey. The cat will slowly walk down forest paths, listening for and stalking prey before rushing or ambushing. The jaguar attacks from cover and usually from a target's blind spot with a quick pounce; the species' ambushing abilities are considered nearly peerless in the animal kingdom by both indigenous people and field researchers and are probably a product of its role as an apex predator in several different environments. The ambush may include leaping into water after prey, as a jaguar is quite capable of carrying a large kill while swimming; its strength is such that carcasses as large as a ] can be hauled up a tree to avoid flood levels. After killing prey, the jaguar will drag the carcass to a ] or other secluded spot. It begins eating at the neck and chest. The heart and lungs are consumed, followed by the shoulders.<ref>{{cite book |chapter=Guidelines for Captive Management of Jaguars |author=Baker, W. K. Jr.|pages=8–16 |editor=Law, C. |title=Jaguar Species Survival Plan |publisher=] |url=http://www.jaguarssp.com/Animal%20Mgmt/JAGUAR%20GUIDELINES.pdf |url-status=dead |archive-url=https://web.archive.org/web/20120113131137/http://www.jaguarssp.com/Animal%20Mgmt/JAGUAR%20GUIDELINES.pdf
|archive-date=13 January 2012}}</ref>

===Social activity===
], ], ], Brazil]]
The jaguar is generally ] except for females with cubs. In 1977, groups consisting of a male, female and cubs, and two females with two males were sighted several times in a study area in the ] valley.<ref name=Schaller1980/> In some areas, males may form paired coalitions which together mark, defend and invade territories, find and mate with the same females and search for and share prey.<ref>{{cite journal|last1=Jędrzejewski|first1=W.|last2=Hoogesteijn|first2=R.|last3=Devlin |first3=A. L. |name-list-style=amp |year=2022 |title=Collaborative behaviour and coalitions in male jaguars (''Panthera onca'')—evidence and comparison with other felids |journal=Behavioral Ecology and Sociobiology |volume=76|issue=9|page=121 |doi=10.1007/s00265-022-03232-3 |s2cid=251713323 |doi-access=free|bibcode=2022BEcoS..76..121J }}</ref> A radio-collared female moved in a ] of {{cvt|25-38|km2}}, which partly overlapped with another female. The home range of the male in this study area overlapped with several females.<ref name=Schaller1980>{{cite journal |author1=Schaller, G. B. |author2=Crawshaw, P. G. Jr. |year=1980 |name-list-style=amp |title=Movement patterns of Jaguar |journal=Biotropica |volume=12 |issue=3 |pages=161–168 |doi=10.2307/2387967 |jstor=2387967|bibcode=1980Biotr..12..161S }}</ref>

The jaguar uses scrape marks, urine, and feces to ].<ref name=Rabinowitz>{{cite journal |author1=Rabinowitz, A. R. |author1-link=Alan Rabinowitz |author2=Nottingham, B.G. Jr. |year=1986 |name-list-style=amp |title=Ecology and behaviour of the Jaguar (''Panthera onca'') in Belize, Central America |journal=Journal of Zoology |volume=210 |issue=1 |pages=149–159 |doi=10.1111/j.1469-7998.1986.tb03627.x}}</ref><ref>{{cite journal |last1=Harmsen |first1=B. J. |last2=Foster |first2=R.J. |last3=Gutierrez |first3=S.M. |last4=Marin |first4=S.Y. |last5=Doncaster |first5=C.P. |year=2007 |name-list-style=amp |title=Scrape-marking behavior of jaguars (''Panthera onca'') and pumas (''Puma concolor'') |journal=Journal of Mammalogy |volume=91 |issue=5 |pages=1225–1234 |doi=10.1644/09-mamm-a-416.1 |doi-access=free}}</ref>
The size of home ranges depends on the level of deforestation and human population density. The home ranges of females vary from {{cvt|15.3|km2}} in the ] to {{cvt|53.6|km2}} in the Amazon to {{cvt|233.5|km2}} in the ]. Male jaguar home ranges vary from {{cvt|25|km2}} in the Pantanal to {{cvt|180.3|km2}} in the Amazon to {{cvt|591.4|km2}} in the Atlantic Forest and {{cvt|807.4|km2}} in the ].<ref>{{cite journal |author1=Morato, R.G. |author2=Stabach, J.A. |author3=Fleming, C.H. |author4=Calabrese, J.M. |author5=De Paula, R.C. |author6=Ferraz, K.M. |author7=Kantek, D.L. |author8=Miyazaki, S.S. |author9=Pereira, T.D. |author10=Araujo, G.R. |author11=Paviolo, A. |year=2016 |name-list-style=amp |title=Space use and movement of a neotropical top predator: the endangered Jaguar |journal=PLOS ONE |volume=11 |issue=12 |page=e0168176 |doi=10.1371/journal.pone.0168176 |doi-access=free |pmc=5193337 |pmid=28030568 |bibcode=2016PLoSO..1168176M}}</ref>
Studies employing ] ] in 2003 and 2004 found densities of only six to seven jaguars per {{cvt|100|km}} in the Pantanal region, compared with 10 to 11 using traditional methods; this suggests the widely used sampling methods may inflate the actual numbers of individuals in a sampling area.<ref name=soisalo/> Fights between males occur but are rare, and avoidance behavior has been observed in the wild.<ref name=Rabinowitz/> In one wetland population with degraded territorial boundaries and more social proximity, adults of the same sex are more tolerant of each other and engage in more friendly and co-operative interactions.<ref name=Eriksson2021/>

]
The jaguar ]s/grunts for long-distance communication;<ref name=Seymour /><ref name=Emmons1987 /> intensive bouts of counter-calling between individuals have been observed in the wild.<ref name=Emmons1987 /> This vocalization is described as "hoarse" with five or six ] notes.<ref name=Seymour /> ] is produced by individuals when greeting, during ], or by a mother comforting her cubs. This sound is described as low intensity snorts, possibly intended to signal tranquility and passivity.<ref>{{Cite journal |last1=Peters |first1=G. |last2=Tonin-Leyhausen |first2=B. |year=1999 |name-list-style=amp |title=Evolution of Acoustic Communication Signals of Mammals: Friendly Close-Range Vocalizations in Felidae (Carnivora) |journal=Journal of Mammalian Evolution |volume=6 |issue=2 |pages=129–159 |doi=10.1023/A:1020620121416 |s2cid=25252052}}</ref><ref>{{cite journal |last1=Leuchtenberger |first1=C. |last2=Crawshaw |first2=P. G. |last3=Mourão |first3=G. |last4=Lehn |first4=C. R. |year=2009 |name-list-style=amp |title=Courtship behavior by Jaguars in the Pantanal of Mato Grosso do Sul |journal=Natureza & Conservação |volume=7 |issue=1 |pages=218–222 |url=https://www.researchgate.net/publication/263424625 |access-date=13 February 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021418/https://www.researchgate.net/publication/263424625_Courtship_behavior_by_Jaguars_in_the_Pantanal_of_Mato_Grosso_do_Sul |url-status=live}}</ref> Cubs have been recorded bleating, gurgling and mewing.<ref name=Seymour />

===Reproduction and life cycle===
] in the northern Pantanal]]
]
In captivity, the female jaguar is recorded to reach ] at the age of about 2.5 years. ] lasts 7–15 days with an ] of 41.8 to 52.6 days. During estrus, she exhibits increased restlessness with rolling and prolonged vocalizations.<ref>{{cite journal |author1=Wildt, D.E. |author2=Platz, C.C. |author3=Chakraborty, P.K. |author4=Seager, S.W.J. |year=1979 |name-list-style=amp |title=Oestrous and ovarian activity in a female jaguar (''Panthera onca'') |journal=Reproduction |volume=56 |issue=2 |pages=555–558 |doi=10.1530/jrf.0.0560555 |doi-access=free |pmid=383976}}</ref>
She is an ] but can also ovulate spontaneously.<ref>{{cite journal |author1=Barnes, S.A. |author2=Teare, J.A. |author3=Staaden, S. |author4=Metrione, L. |author5=Penfold, L.M. |year=2016 |name-list-style=amp |title=Characterization and manipulation of reproductive cycles in the jaguar (''Panthera onca'') |journal=General and Comparative Endocrinology |volume=225 |pages=95–103 |doi=10.1016/j.ygcen.2015.09.012 |doi-access=free |pmid=26399935}}</ref><ref>{{cite journal |author1=Jorge-Neto, P. N. |name-list-style=amp |author2=Luczinski, T. C. |author3=Ribeiro de Araújo, G. |author4=Salomão, J. |author5=de Souza Traldi, A. |author6=Melo dos Santos, J. A. |author7=Requena, L. A. |author8=Machado Gianni, M. C. |author9= de Deco-Souza, T. |author10=Schilbach Pizzutto, C. |author11=Baldassarre, H. |year=2020 |journal=Theriogenology |volume=147 |pages=57–61 |title=Can jaguar (''Panthera onca'') ovulate without copulation? |doi=10.1016/j.theriogenology.2020.02.026|doi-access=free |pmid=32092606 }}</ref>
] lasts 91 to 111 days.<ref>{{cite book |author=Hemmer, H. |year=1976 |chapter=Gestation period and postnatal development in felids |title=The world's cats |editor=Eaton, R.L. |publisher=Carnivore Research Institute, Univ. Washington |location=Seattle |volume=3. Contributions to Biology, Ecology, Behavior and Evolution |pages=143–165}}</ref>
The male is sexually mature at the age of three to four years.<ref>{{cite book |author1=Mondolfi, E. |author2=Hoogesteijn, R. |year=1986 |name-list-style=amp |chapter=Notes on the biology and status of the jaguar (''Panthera onca'') in Venezuela |title=Cats of the world: biology, conservation and management |editor1=Miller, S.D. |editor2=Everett, D.D. |publisher=National Wildlife Federation |location=Washington, DC |pages=85–123 |isbn=978-091218679-5}}</ref>
His mean ] volume is 8.6±1.3&nbsp;ml.<ref>{{cite journal |author1=Morato, R. G. |author2=Guimaraes, M.A.B. |author3=Ferriera, F. |author4=Verreschi, I.T.d.N. |author5=Barnabe, R.C. |year=1999 |name-list-style=amp |title=Reproductive characteristics of captive male jaguars |journal=Brazilian Journal of Veterinary Research and Animal Science |volume=36 |issue=5 |doi=10.1590/S1413-95961999000500008 |doi-access=free}}</ref>
] of the jaguar is 9.8 years.<ref>{{cite journal |author1=Pacifici, M. |author2=Santini, L. |author3=Di Marco, M. |author4=Baisero, D. |author5=Francucci, L. |author6=Grottolo Marasini, G. |author7=Visconti, P. |author8=Rondinini, C. |name-list-style=amp |year=2013 |title=Generation length for mammals |journal=Nature Conservation |volume=5 |issue=5 |pages=87–94 |doi=10.3897/natureconservation.5.5734 |doi-access=free}}</ref>

In the Pantanal, ]s were observed to stay together for up to five days. Females had one to two cubs.<ref>{{cite journal |author1=Cavalcanti, S. M. C. |author2=Gese, E. M. |year=2009 |name-list-style=amp |title=Spatial ecology and social interactions of jaguars (''Panthera onca'') in the southern Pantanal, Brazil |journal=Journal of Mammalogy |volume=90 |issue=4 |pages=935–945 |doi=10.1644/08-MAMM-A-188.1 |doi-access=free}}</ref>
The young are born with closed eyes but open them after two weeks. Cubs are ]ed at the age of three months but remain in the birth den for six months before leaving to accompany their mother on hunts.<ref name=SWWL>{{cite magazine |author=Egerton, J. |date=2006 |title=Jaguars: Magnificence in the Southwest |magazine=Wild Tracks |url=http://www.southwestwildlife.org/pdf/Newsletter/Spring06.pdf |url-status=dead |archive-url=https://web.archive.org/web/20110721144027/http://www.southwestwildlife.org/pdf/Newsletter/Spring06.pdf |access-date=6 December 2009 |archive-date=21 July 2011}}</ref>
Jaguars remain with their mothers for up to two years. They appear to rarely live beyond 11 years, but captive individuals may live 22 years.<ref name=Seymour />

In 2001, a male jaguar killed and partially consumed two cubs in ]. ] of blood samples revealed that the male was the father of the cubs.<ref>{{cite journal |author1=Soares, T. N. |author2=Telles, M. P. |author3=Resende, L.V. |author4=Silveira, L. |author5=Jácomo, A.T.A. |author6=Morato, R.G. |author7=Diniz-Filho, J.A.F. |author8=Eizirik, E. |author9=Brondani, R.P. |author10=Brondani, C. |year=2006 |display-authors=4 |name-list-style=amp |title=Paternity testing and behavioral ecology: A case study of jaguars (''Panthera onca'') in Emas National Park, Central Brazil |journal=Genetics and Molecular Biology |volume=29 |issue=4 |pages=735–740 |doi=10.1590/S1415-47572006000400025 |doi-access=free}}</ref> Two more cases of ] were documented in the northern Pantanal in 2013.<ref>{{cite journal |author1=Tortato, F.R. |author2=Devlin, A.L. |author3=Hoogesteijn, R. |author4=Júnior, J.A.M. |author5=Frair, J.L. |author6=Crawshaw, P.G. |author7=Izzo, T.J. |author8=Quigley, H.B. |year=2017 |name-list-style=amp |title=Infanticide in a jaguar (''Panthera onca'') population – does the provision of livestock carcasses increase the risk? |journal=Acta Ethologica |volume=20 |issue=1 |pages=69–73 |doi=10.1007/s10211-016-0241-4 |s2cid=34002056 |url=https://www.researchgate.net/publication/308940647 |access-date=27 February 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021302/https://www.researchgate.net/publication/308940647_Infanticide_in_a_jaguar_Panthera_onca_population-does_the_provision_of_livestock_carcasses_increase_the_risk |url-status=live}}</ref> To defend against infanticide, the female may hide her cubs and distract the male with courtship behavior.<ref>{{cite journal |author1=Stasiukynas, D. C. |author2=Boron, V. |author3=Hoogesteijn, R. |author4=Barragán2, J. |author5=Martin, A. |author6=Tortato, F. |author7= Rincón, S. |author8=Payán, E. |year=2021 |title=Hide and flirt: observed behavior of female jaguars (''Panthera onca'') to protect their young cubs from adult males |journal=Acta Ethologica |volume=25 |issue=3 |pages=179–183 |doi=10.1007/s10211-021-00384-9|s2cid=239539707}}</ref>

===Attacks on humans===
{{Further|Man-eating animal#Jaguars}}
The Spanish ]s feared the jaguar. According to ], the ] stated that people did not need to fear the jaguar as long as capybaras were abundant.<ref name=Porter1894>{{cite book |author=Porter, J. H. |year=1894 |chapter=The Jaguar |title=Wild beasts; a study of the characters and habits of the elephant, lion, leopard, panther, jaguar, tiger, puma, wolf, and grizzly bear |publisher=C. Scribner's sons |location=New York |chapter-url=https://archive.org/stream/wildbeastsstud00port#page/n197/mode/2up |pages=174–195}}</ref>
The first official record of a jaguar killing a human in Brazil dates to June 2008.<ref>{{cite journal |author1=De Paula, R. |author2=Campos Neto, M. F. |author3=Morato, R. G. |year=2008 |name-list-style=amp |title=First official record of Human killed by Jaguar in Brazil |journal=Cat News |issue=49 |pages=31–32}}</ref>
Two children were attacked by jaguars in Guyana.<ref name=r22>{{cite journal |author1=Iserson, K. V. |author2=Francis, A. M. |year=2015 |name-list-style=amp |title=Jaguar attack on a child: Case report and literature review |journal=Western Journal of Emergency Medicine |volume=16 |issue=2 |pages=303–309 |doi=10.5811/westjem.2015.1.24043 |pmc=4380383 |pmid=25834674}}</ref>
The majority of known attacks on people happened when it had been cornered or wounded.<ref>{{Cite book |last1=Seidensticker |first1=J. |last2=Lumpkin |first2=S. |date=2016 |name-list-style=amp |title=Cats in Question: The Smithsonian Answer Book |publisher=Smithsonian Institution |location=Washington, D.C. |url=https://books.google.com/books?id=09LwCwAAQBAJ&pg=PT303 |isbn=978-158834546-2}}</ref>

==Threats==
]]]
The jaguar is threatened by ] and ] of habitat, illegal killing in retaliation for livestock depredation and for illegal trade in jaguar body parts. It is listed as ] on the ] since 2002, as the jaguar population has probably declined by 20–25% since the mid-1990s. ] is a major threat to the jaguar across its range. Habitat loss was most rapid in drier regions such as the Argentine ], the arid grasslands of Mexico and the southwestern United States.<ref name=iucn/>

In 2002, it was estimated that the range of the jaguar had declined to about 46% of its range in the early 20th century.<ref name=Sanderson2002/> In 2018, it was estimated that its range had declined by 55% in the last century. The only remaining stronghold is the Amazon rainforest, a region that is rapidly being fragmented by deforestation.<ref>{{cite journal |author1=De La Torre, J.A. |author2=González-Maya, J.F. |author3=Zarza, H. |author4=Ceballos, G. |author5=Medellín, R.A. |year=2018 |name-list-style=amp |title=The jaguar's spots are darker than they appear: assessing the global conservation status of the jaguar ''Panthera onca'' |journal=Oryx |volume=52 |issue=2 |pages=300–315 |doi=10.1017/S0030605316001046 |doi-access=free}}</ref>
Between 2000 and 2012, forest loss in the jaguar range amounted to {{convert|83.759|km2|abbr=on}}, with fragmentation increasing in particular in corridors between Jaguar Conservation Units (JCUs).<ref>{{cite journal |author1=Olsoy, P.J. |author2=Zeller, K.A. |author3=Hicke, J.A. |author4=Quigley, H.B. |author5=Rabinowitz, A.R. |author6=Thornton, D.H. |year=2016 |name-list-style=amp |title=Quantifying the effects of deforestation and fragmentation on a range-wide conservation plan for jaguars |journal=Biological Conservation |volume=203 |pages=8–16 |url=https://www.researchgate.net/publication/307954092 |doi=10.1016/j.biocon.2016.08.037 |bibcode=2016BCons.203....8O |access-date=13 April 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021307/https://www.researchgate.net/publication/307954092_Quantifying_the_effects_of_deforestation_and_fragmentation_on_a_range-wide_conservation_plan_for_jaguars |url-status=live}}</ref>
By 2014, direct linkages between two JCUs in Bolivia were lost, and two JCUs in northern Argentina became completely isolated due to deforestation.<ref>{{cite journal |author1=Thompson, J.J. |author2=Velilla, M. |year=2017 |name-list-style=amp |title=Modeling the effects of deforestation on the connectivity of jaguar ''Panthera onca'' populations at the southern extent of the species' range |journal=Endangered Species Research |volume=34 |pages=109–121 |url=https://www.int-res.com/articles/esr2017/34/n034p109.pdf |doi=10.3354/esr00840 |doi-access=free |access-date=2 March 2021 |archive-date=8 March 2021 |archive-url=https://web.archive.org/web/20210308132647/https://www.int-res.com/articles/esr2017/34/n034p109.pdf |url-status=live}}</ref>

In Mexico, the jaguar is primarily threatened by ]. Its habitat is fragmented in northern Mexico, in the ] and the ], caused by changes in land use, construction of roads and tourism infrastructure.<ref name=Ceballos2016>{{cite book |author1=Ceballos, G. |author2=Zarza, H. |author3=Chávez, C. |author4=González-Maya, J.F. |year=2016 |name-list-style=amp |chapter=Ecology and Conservation of Jaguars in Mexico |title=Tropical conservation: Perspectives on local and global priorities |editor1=Aguirre, A. |editor2=Sukumar, R. |publisher=Oxford University Press |chapter-url=https://www.researchgate.net/publication/307985045 |pages=273–289 |isbn=978-019976698-7 |access-date=28 February 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021317/https://www.researchgate.net/publication/307985045_Ecology_and_conservation_of_jaguars_in_Mexico_state_of_knowledge_and_future_challenges |url-status=live}}</ref>
In Panama, 220 of 230 jaguars were killed in retaliation for predation on livestock between 1998 and 2014.<ref>{{cite journal |author1=Moreno, R. |author2=Meyer, N. |author3=Olmos, M. |author4=Hoogesteijn, R. |author5=Hoogesteijn, A.L. |year=2015 |name-list-style=amp |title=Causes of jaguar killing in Panama – a long term survey using interviews |journal=Cat News |issue=62 |pages=40–42 |url=https://repository.si.edu/bitstream/handle/10088/29640/CN62_Moreno_et_al_2015.pdf |access-date=18 November 2021 |archive-date=18 November 2021 |archive-url=https://web.archive.org/web/20211118072049/https://repository.si.edu/bitstream/handle/10088/29640/CN62_Moreno_et_al_2015.pdf |url-status=live}}</ref>
In Venezuela, the jaguar was extirpated in about 26% of its range in the country since 1940, mostly in dry ]s and unproductive scrubland in the northeastern region of ].<ref>{{cite journal |last1=Jędrzejewski |first1=W. |last2=Boede |first2=E.O. |last3=Abarca |first3=M. |last4=Sánchez-Mercado |first4=A. |last5=Ferrer-Paris |first5=J.R. |last6=Lampo |first6=M. |last7=Velásquez |first7=G. |last8=Carreño |first8=R. |last9=Viloria |first9=Á.L. |last10=Hoogesteijn |first10=R. |last11=Robinson |first11=H.S. |last12=Stachowicz |first12=I. |last13=Cerda |first13=H. |last14=Weisz |first14=M. del Mar |last15=Barros |first15=T.R. |last16=Rivas |first16=Gilson A. |last17=Borges |first17=G. |last18=Molinari |first18=J. |last19=Lew |first19=D. |last20=Takiff |first20=H. |last21=Schmidt |first21=K. |year=2017 |name-list-style=amp |title=Predicting carnivore distribution and extirpation rate based on human impacts and productivity factors; assessment of the state of jaguar (''Panthera onca'') in Venezuela |journal=Biological Conservation |volume=206 |pages=132–142 |url=https://www.researchgate.net/publication/312059394 |doi=10.1016/j.biocon.2016.09.027 |bibcode=2017BCons.206..132J |access-date=28 February 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021303/https://www.researchgate.net/publication/312059394_Predicting_carnivore_distribution_and_extirpation_rate_based_on_human_impacts_and_productivity_factors_assessment_of_the_state_of_jaguar_Panthera_onca_in_Venezuela |url-status=live}}</ref>
In Ecuador, the jaguar is threatened by reduced prey availability in areas where the expansion of the road network facilitated access of human hunters to forests.<ref>{{cite journal |author1=Espinosa, S. |author2=Celis, G. |author3=Branch, L.C. |year=2018 |name-list-style=amp |title=When roads appear jaguars decline: Increased access to an Amazonian wilderness area reduces potential for jaguar conservation |journal=PLOS ONE |volume=13 |issue=1 |page=e0189740 |doi=10.1371/journal.pone.0189740 |doi-access=free |pmc=5751993 |pmid=29298311 |bibcode=2018PLoSO..1389740E}}</ref>
In the ], at least 117 jaguars were killed in ] and the adjacent ] between 1995 and 2008.<ref>{{cite journal |author1=Paviolo, A. |author2=De Angelo, C.D. |author3=Di Blanco, Y.E. |author4=Di Bitetti, M.S. |year=2008 |name-list-style=amp |title=Jaguar ''Panthera onca'' population decline in the Upper Paraná Atlantic Forest of Argentina and Brazil |journal=Oryx |volume=42 |issue=4 |pages=554–561 |doi=10.1017/S0030605308000641 |doi-broken-date=1 November 2024 |doi-access=free|hdl=11336/61266 |hdl-access=free}}</ref>
Some ] in the Colombian ] hunt jaguars for consumption and sale of meat.<ref>{{cite journal |author1=Balaguera-Reina, S. |author2=Gonzalez-Maya, J.F. |year=2008 |name-list-style=amp |title=Occasional jaguar hunting for subsistence in Colombian Chocó |journal=Cat News |issue=48 |page=5 |url=https://www.researchgate.net/publication/233399315 |access-date=28 February 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021347/https://www.researchgate.net/publication/233399315_Occasional_Jaguar_Hunting_for_Subsistence_in_Colombian_Choco |url-status=live}}</ref>
Between 2008 and 2012, at least 15 jaguars were killed by livestock farmers in central Belize.<ref>{{cite journal |author1=Foster, R.J. |author2=Harmsen, B.J. |author3=Urbina, Y. L. |author4=Wooldridge, R.L. |author5=Doncaster, C.P. |author6=Quigley, H. |author7=Figueroa, O.A. |year=2020 |name-list-style=amp |title=Jaguar (''Panthera onca'') density and tenure in a critical biological corridor |journal=Journal of Mammalogy |volume=101 |issue=6 |pages=1622–1637 |doi=10.1093/jmammal/gyaa134 |pmid=33505226 |pmc=7816682 |doi-access=free}}</ref>

The international trade of jaguar skins boomed between the end of the ] and the early 1970s.<ref name="SkinTrade">{{cite book |last=Broad |first=S. |date=1987 |title=The harvest of and trade in Latin American spotted cats (Felidae) and otters (Lutrinae) |publisher=IUCN Conservation Monitoring Centre |location=Cambridge |url=https://www.biodiversitylibrary.org/item/119261 |access-date=21 February 2018 |archive-date=13 January 2020 |archive-url=https://web.archive.org/web/20200113095346/https://www.biodiversitylibrary.org/item/119261 |url-status=live}}</ref>
Significant declines occurred in the 1960s, as more than 15,000 jaguars were yearly killed for their skins in the ] alone; the trade in jaguar skins decreased since 1973 when the ] was enacted.<ref name="r32">{{cite journal |last1=Weber |first1=W. |author2=Rabinowitz, A. |year=1996 |name-list-style=amp |title=A global perspective on large carnivore conservation |journal=Conservation Biology |volume=10 |issue=4 |pages=1046–1054 |doi=10.1046/j.1523-1739.1996.10041046.x |bibcode=1996ConBi..10.1046W |url=http://www.jaguarnetwork.org/pdf/71.pdf |archive-url=https://web.archive.org/web/20120426042409/http://www.jaguarnetwork.org/pdf/71.pdf |archive-date=26 April 2012}}</ref>
Interview surveys with 533 people in the northwestern Bolivian Amazon revealed that local people killed jaguars out of fear, in retaliation, and for trade.<ref>{{cite journal |author1=Knox, J. |author2=Negrões, N. |author3=Marchini, S. |author4=Barboza, K. |author5=Guanacoma, G. |author6=Balhau, P. |author7=Tobler, M.W. |author8=Glikman, J.A. |year=2019 |name-list-style=amp |title=Jaguar persecution without "cowflict": insights from protected territories in the Bolivian Amazon |journal=Frontiers in Ecology and Evolution |volume=7 |page=494 |doi=10.3389/fevo.2019.00494 |doi-access=free}}</ref>
Between August 2016 and August 2019, jaguar skins and body parts were seen for sale in tourist markets in the Peruvian cities of ], ] and ].<ref>{{cite journal |author1=Braczkowski, A. |author2=Ruzo, A. |author3=Sanchez, F. |author4=Castagnino, R. |author5=Brown, C. |author6=Guynup, S. |author7=Winter, S. |author8=Gandy, D. |author9=O'Bryan, C. |year=2019 |name-list-style=amp |title=The ayahuasca tourism boom: An undervalued demand driver for jaguar body parts? |journal=Conservation Science and Practice |volume=1 |issue=12 |page=e126 |url=https://www.researchgate.net/publication/336450042 |doi=10.1111/csp2.126 |doi-access=free |bibcode=2019ConSP...1E.126B |access-date=28 February 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021318/https://www.researchgate.net/publication/336450042_The_ayahuasca_tourism_boom_An_undervalued_demand_driver_for_jaguar_body_parts |url-status=live }}</ref>
], opportunistic hunting and hunting for trade in domestic markets are key drivers for killing jaguars in Belize and Guatemala.<ref>{{cite journal |author1=Arias, M. |author2=Hinsley, A. |author3=Milner-Gulland, E.J. |year=2020 |name-list-style=amp |title=Characteristics of, and uncertainties about, illegal jaguar trade in Belize and Guatemala |journal=Biological Conservation |volume=250 |page=108765 |doi=10.1016/j.biocon.2020.108765 |bibcode=2020BCons.25008765A |s2cid=224967913 |url=https://www.researchgate.net/publication/344172628 |access-date=18 November 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021310/https://www.researchgate.net/publication/344172628_Characteristics_of_and_uncertainties_about_illegal_jaguar_trade_in_Belize_and_Guatemala |url-status=live}}</ref>
Seizure reports indicate that at least 857 jaguars were involved in trade between 2012 and 2018, including 482 individuals in Bolivia alone; 31 jaguars were seized in ].<ref>{{cite journal |author1=Morcatty, T.Q. |author2=Bausch Macedo, J.C. |author3=Nekaris, K.A.I. |author4=Ni, Q. |author5=Durigan, C.C. |author6=Svensson, M.S. |author7=Nijman, V. |year=2020 |name-list-style=amp |title=Illegal trade in wild cats and its link to Chinese-led development in Central and South America |journal=Conservation Biology |volume=34 |issue=6 |pages=1525–1535 |doi=10.1111/cobi.13498 |pmid=32484587 |doi-access=free|bibcode=2020ConBi..34.1525M}}</ref>
Between 2014 and early 2019, 760 jaguar fangs were seized that originated in Bolivia and were destined for China. Undercover investigations revealed that the ] of jaguar body parts is run by Chinese residents in Bolivia.<ref>{{cite report |author=Earth League International |year=2020 |title=Unveiling the criminal networks behind jaguar trafficking in Bolivia |publisher=IUCN National Committee of the Netherlands |location=Amsterdam |url=https://www.iucn.nl/app/uploads/2021/03/iucn_nl_report_jaguar_trafficking_bolivia_media-1.pdf |access-date=26 July 2021 |archive-date=26 July 2021 |archive-url=https://web.archive.org/web/20210726070809/https://www.iucn.nl/app/uploads/2021/03/iucn_nl_report_jaguar_trafficking_bolivia_media-1.pdf |url-status=live}}</ref>

==Conservation==
The jaguar is listed on ], which means that all international commercial trade in jaguars or their body parts is prohibited. Hunting jaguars is prohibited in Argentina, Brazil, Colombia, French Guiana, Honduras, Nicaragua, Panama, Paraguay, Suriname, the United States, and Venezuela. Hunting jaguars is restricted in Guatemala and Peru.<ref name=iucn /> In Ecuador, hunting jaguars is prohibited, and it is classified as threatened with extinction.<ref>{{cite book |author1=Zapata Ríos, G. |author2=Araguillin, E. |author3=Cevallos, J. |author4=Moreno, F. |author5=Ortega, A. |author6=Rengel, J. |author7=Valarezo, N. |date=2014 |name-list-style=amp |title=Plan de Acción para la Conservación del Jaguar en el Ecuador |trans-title=Action Plan for the Conservation of the Jaguar in Ecuador |publisher=Ministerio del Ambiente y Wildlife Conservation Society Ecuador |location=Quito |language=es |url=http://www.wild4ever.com/wp-content/uploads/2012/05/Ecuador-National-Jaguar-Plan.pdf |access-date=13 February 2021 |archive-date=23 September 2020 |archive-url=https://web.archive.org/web/20200923073232/http://www.wild4ever.com/wp-content/uploads/2012/05/Ecuador-National-Jaguar-Plan.pdf |url-status=live }}</ref>
In Guyana, it is protected as an endangered species, and hunting it is illegal.<ref>{{cite book |author=Kerman, I. |date=2010 |title=Exploitation of the jaguar, ''Panthera onca'' and other large forest cats in Suriname |editor=Felix, M.-L. |publisher=WWF Guianas |location=Paramaribo |url=https://www.a2000greetings.com/downloads/exploitation_of_the_jaguar_and_other_large_forest_cats_in_suriname_irvin_kerman.pdf |access-date=3 March 2021 |archive-date=18 September 2021 |archive-url=https://web.archive.org/web/20210918010124/https://www.a2000greetings.com/downloads/exploitation_of_the_jaguar_and_other_large_forest_cats_in_suriname_irvin_kerman.pdf |url-status=live }}</ref>
], a jaguar in Arizona]]
In 1986, the ] was established in Belize as the world's first protected area for jaguar conservation.<ref name=Weckel>{{cite journal |author1=Weckel, M. |author2=Giuliano, W. |author3=Silver, S. |year=2006 |name-list-style=amp |title=Cockscomb revisited: jaguar diet in the Cockscomb Basin Wildlife Sanctuary, Belize |journal=Biotropica |volume=38 |issue=5 |pages=687–690 |doi=10.1111/j.1744-7429.2006.00190.x |bibcode=2006Biotr..38..687W |s2cid=85151201 }}</ref>

===Jaguar Conservation Units===
In 1999, field scientists from 18 jaguar range countries determined the most important areas for long-term jaguar conservation based on the status of jaguar population units, stability of prey base and quality of habitat. These areas, called "Jaguar Conservation Units" (JCUs), are large enough for at least 50 breeding individuals and range in size from {{cvt|566|to|67598|km2}}; 51 JCUs were designated in 36 geographic regions including:<ref name=Sanderson2002 />
* the ] and ] in Mexico
* the ] tropical forests extending over Mexico, Belize and Guatemala
* the ] from Honduras and Panama to Colombia
* ]
* northern ] and ] in Brazil
* ] in Bolivia and Peru
* ] in Argentina

Optimal routes of travel between core jaguar population units were identified across its range in 2010 to implement ]s that connect JCUs. These corridors represent areas with the shortest distance between jaguar breeding populations, require the least possible energy input of ] individuals and pose a low mortality risk. They cover an area of {{cvt|2600000|km2}} and range in length from {{cvt|3|to|1102|km}} in Mexico and Central America and from {{cvt|489.14|to|1607|km}} in South America.<ref name=RabinowitzZeller2010>{{Cite journal |author1=Rabinowitz, A. |author2=Zeller, K.A. |year=2010 |name-list-style=amp |title=A range-wide model of landscape connectivity and conservation for the jaguar, ''Panthera onca'' |journal=Biological Conservation |volume=143 |issue=4 |pages=939–945 |doi=10.1016/j.biocon.2010.01.002 |bibcode=2010BCons.143..939R |url=https://www.panthera.org/cms/sites/default/files/documents/Rabinowitz_Zeller_2010_Arangewidemodeloflandscapeconnectivityandconservationforjaguar_BioCon.pdf |access-date=14 March 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021304/https://www.panthera.org/cms/sites/default/files/documents/Rabinowitz_Zeller_2010_Arangewidemodeloflandscapeconnectivityandconservationforjaguar_BioCon.pdf |url-status=live }}</ref>
Cooperation with local landowners and municipal, state, or federal agencies is essential to maintain connected populations and prevent fragmentation in both JCUs and corridors.<ref>{{cite book |author1=Zeller, K.A. |author2=Rabinowitz, A. |author3=Salom-Perez, R. |author4=Quigley, H. |year=2013 |name-list-style=amp |chapter=The Jaguar Corridor Initiative: A range-wide conservation strategy |title=Molecular population genetics, evolutionary biology and biological conservation of Neotropical carnivores |editor1=Ruiz-Garcia, M. |editor2=Shostell, J.M. |publisher=Nova Science Publishers |location=New York |chapter-url=https://conservationcorridor.org/cpb/Zeller_et_al_2013.pdf |pages=629–657 |isbn=978-1-62417-071-3 |access-date=14 March 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129021304/https://conservationcorridor.org/cpb/Zeller_et_al_2013.pdf |url-status=live }}</ref>
Seven of 13 corridors in Mexico are functioning with a width of at least {{cvt|14.25|km}} and a length of no more than {{cvt|320|km}}. The other corridors may hamper passage, as they are narrower and longer.<ref>{{cite journal |author1=Rodríguez-Soto, C. |author2=Monroy-Vilchis, O. |author3=Zarco-González, M.M. |year=2013 |name-list-style=amp |title=Corridors for jaguar (''Panthera onca'') in Mexico: Conservation strategies |journal=Journal for Nature Conservation |volume=21 |issue=6 |pages=438–443 |doi=10.1016/j.jnc.2013.07.002 |bibcode=2013JNatC..21..438R |url=https://www.academia.edu/35225702 |access-date=18 November 2021 |archive-date=29 November 2021 |archive-url=https://web.archive.org/web/20211129044432/https://www.academia.edu/35225702 |url-status=live }}</ref>

In August 2012, the ] set aside {{cvt|838232|acres|km2|order=flip}} in Arizona and New Mexico for the protection of the jaguar.<ref>{{cite journal |author=Department of the Interior Fish and Wildlife Service |year=2012 |title=Designation of Critical Habitat for Jaguar; Proposed Rule |journal=Federal Register |volume=77 |issue=161 |pages=50214–50242 |url=https://www.fws.gov/southwest/es/arizona/Documents/SpeciesDocs/Jaguar/Jaguar_pCH_FR_8-20-2012.pdf |access-date=13 March 2021 |archive-date=14 February 2020 |archive-url=https://web.archive.org/web/20200214001827/https://www.fws.gov/southwest/es/arizona/Documents/SpeciesDocs/Jaguar/Jaguar_pCH_FR_8-20-2012.pdf |url-status=live }}</ref> The Jaguar Recovery Plan was published in April 2019, in which ] is considered to form the northern boundary of the Jaguar Recovery Unit in Arizona and New Mexico.<ref>{{cite journal |author1=Sanderson, E.W. |name-list-style=amp |author2=Fisher, K. |author3=Peters, R. |author4=Beckmann, J.P. |author5=Bird, B. |author6=Bradley, C.M. |author7=Bravo, J.C. |author8=Grigione, M.M. |author9=Hatten, J.R. |author10=González, C.A.L. |author11=Menke, K. |author12=Miller, J.R.B. |author13=Miller, P.S. |author14=Mormorunni, C. |author15=Robinson, M.J. |author16=Thomas, R.E. |author17=Wilcox, S. |year=2021 |title=A systematic review of potential habitat suitability for the jaguar ''Panthera onca'' in central Arizona and New Mexico, USA |journal=Oryx |volume= 56|issue= |pages=116–127 |doi=10.1017/S0030605320000459 |doi-access=free}}</ref>

In Mexico, a national conservation strategy was developed from 2005 on and published in 2016.<ref name=Ceballos2016/> The Mexican jaguar population increased from an estimated 4,000 individuals in 2010 to about 4,800 individuals in 2018. This increase is seen as a positive effect of conservation measures that were implemented in cooperation with governmental and non-governmental institutions and landowners.<ref>{{cite journal |author1=Ceballos, G. |name-list-style=amp |author2=Zarza, H. |author3=González-Maya, J.F. |author4=de la Torre, J.A. |author5=Arias-Alzate, A. |author6=Alcerreca, C. |author7=Barcenas, H.V. |author8=Carreón-Arroyo, G. |author9=Chávez, C. |author10=Cruz, C. |author11=Medellín, D. |year=2021 |title=Beyond words: From jaguar population trends to conservation and public policy in Mexico |journal=PLOS ONE |volume=16 |issue=10 |pages=e0255555 |doi=10.1371/journal.pone.0255555 |pmid=34613994 |pmc=8494370 |bibcode=2021PLoSO..1655555C |doi-access=free}}</ref>

An evaluation of JCUs from Mexico to Argentina revealed that they overlap with high-quality habitats of about 1,500 mammals to varying degrees. Since co-occurring mammals benefit from the JCU approach, the jaguar has been called an ].<ref>{{cite journal |author1=Thornton, D. |name-list-style=amp |author2=Zeller, K. |author3=Rondinini, C. |author4=Boitani, L. |author5=Crooks, K. |author6=Burdett, C. |author7=Rabinowitz, A. |author8=Quigley, H. |year=2016 |title=Assessing the umbrella value of a range-wide conservation network for jaguars (''Panthera onca'') |journal=Ecological Applications |volume=26 |issue=4 |pages=1112–1124 |doi=10.1890/15-0602 |jstor=24818150 |pmid=27509752 |bibcode=2016EcoAp..26.1112T |hdl=11573/893793 |url=https://iris.uniroma1.it/retrieve/handle/11573/893793/279408/Thornton_Assessing_2016.pdf |access-date=16 November 2021 |archive-date=16 November 2021 |archive-url=https://web.archive.org/web/20211116083400/https://iris.uniroma1.it/retrieve/handle/11573/893793/279408/Thornton_Assessing_2016.pdf |url-status=live }}</ref>
Central American JCUs overlap with the habitat of 187 of 304 regional endemic amphibian and reptile species, of which 19 amphibians occur only in the jaguar range.<ref>{{cite journal |author1=Figel, J.J. |name-list-style=amp |author2=Castañeda, F. |author3=Calderón, A.P. |author4=Torre, J. |author5=García-Padilla, E. |author6=Noss, R.F. |year=2018 |title=Threatened amphibians sheltered under the big cat's umbrella: conservation of jaguars ''Panthera onca'' (Carnivora: Felidae) and endemic herpetofauna in Central America |journal=Revista de Biología Tropical |volume=66 |issue=4 |pages=1741–1753 |doi=10.15517/rbt.v66i4.32544 |doi-access=free}}</ref>

===Approaches===
]]]
In setting up protected reserves, efforts generally also have to be focused on the surrounding areas, as jaguars are unlikely to confine themselves to the bounds of a reservation, especially if the population is increasing in size. Human attitudes in the areas surrounding reserves and laws and regulations to prevent poaching are essential to make conservation areas effective.<ref name=gutierrez>{{cite journal |last1=Gutierrez-Gonzalez |first1=C.E. |last2=Gomez-Ramirez |first2=M.A. |last3=Lopez-Gonzalez |first3=C.A. |last4=Doherty |first4=P.F. |year=2015 |name-list-style=amp |title=Are Private Reserves Effective for Jaguar Conservation? |journal=PLOS ONE |volume=10 |issue=9 |page=e0137541 |bibcode=2015PLoSO..1037541G |doi=10.1371/journal.pone.0137541 |pmc=4580466 |pmid=26398115 |doi-access=free}}</ref>

To estimate population sizes within specific areas and to keep track of individual jaguars, ]ping and ] are widely used, and feces are sought out with the help of ]s to study jaguar health and diet.<ref name=soisalo>{{cite journal |last1=Soisalo |first1=M.K. |last2=Cavalcanti |first2=S.M.C. |year=2006 |name-list-style=amp |title=Estimating the density of a Jaguar population in the Brazilian Pantanal using camera-traps and capture-recapture sampling in combination with GPS radio-telemetry |journal=Biological Conservation |volume=129 |issue=4 |pages=487–496 |doi=10.1016/j.biocon.2005.11.023 |bibcode=2006BCons.129..487S |url=http://www.ekonoiz.com/Eco_Projects/Jaguar_Conservation/estimatingthedensityofjaguarsinthepantanal.pdf |access-date=29 November 2021 |archive-date=13 January 2022 |archive-url=https://web.archive.org/web/20220113193618/http://www.ekonoiz.com/Eco_Projects/Jaguar_Conservation/estimatingthedensityofjaguarsinthepantanal.pdf |url-status=live }}</ref><ref name=furtado>{{cite journal |last1=Furtado |first1=M.M. |last2=Carrillo-Percastegui |first2=S.E. |last3=Jácomo |first3=A.T.A. |last4=Powell |first4=G. |last5=Silveira |first5=L. |last6=Vynne |first6=C. |last7=Sollmann |first7=R. |year=2008 |name-list-style=amp |title=Studying jaguars in the wild: past experiences and future perspectives |journal=Cat News |issue=Special Issue 4 |pages=41–47 |url=http://www.catsg.org/fileadmin/filesharing/5.Cat_News/5.3._Special_Issues/5.3.4._SI_4/Furtado_et_al_2008_Jaguar_field_methods_s.pdf |access-date=2 February 2019 |archive-date=2 February 2019 |archive-url=https://web.archive.org/web/20190202212055/http://www.catsg.org/fileadmin/filesharing/5.Cat_News/5.3._Special_Issues/5.3.4._SI_4/Furtado_et_al_2008_Jaguar_field_methods_s.pdf |url-status=live}}</ref>

Current conservation efforts often focus on educating ranch owners and promoting ].<ref name=WWF>{{cite web |author=Estévez, E. |date=2009 |title=Jaguar Refuge in the Llanos Ecoregion |publisher=] |url=http://wwf.panda.org/es/nuestro_trabajo/latinoamerica/venezuela/index.cfm?uProjectID=VE0854 |url-status=dead |archive-url=https://web.archive.org/web/20141217001905/http://wwf.panda.org/es/nuestro_trabajo/latinoamerica/venezuela/index.cfm?uProjectID=VE0854 |access-date= 1 September 2006 |archive-date=17 December 2014}}</ref> Ecotourism setups are being used to generate public interest in charismatic animals such as the jaguar while at the same time generating revenue that can be used in conservation efforts. A key concern in jaguar ecotourism is the considerable habitat space the species requires. If ecotourism is used to aid in jaguar conservation, some considerations need to be made as to how existing ecosystems will be kept intact, or how new ecosystems will be put into place that are large enough to support a growing jaguar population.<ref name=mossaz>{{cite journal |last1=Mossaz |first1=A. |last2=Buckley |first2=R.C. |last3=Castley |first3=J.G. |year=2015 |name-list-style=amp |title=Ecotourism contributions to conservation of African big cats |journal=Journal for Nature Conservation |volume=28 |pages=112–118 |doi=10.1016/j.jnc.2015.09.009 |bibcode=2015JNatC..28..112M |hdl=10072/125191 |hdl-access=free}}</ref>

Conservationists and professionals in Mexico and the United States have established the {{cvt|56,000|acre}} ] in northern Mexico. Advocacy for reintroduction of the jaguar to its former range in Arizona and New Mexico have been supported by documentation of natural migrations by individual jaguars into the southern reaches of both states, the recency of extirpation from those regions by human action, and supportive arguments pertaining to biodiversity, ecological, human, and practical considerations.<ref>{{cite journal |last1=Sanderson |first1=E. W. |author2=Beckmann, J. P. |author3=Beier, P. |author4=Bird, B. |author5=Bravo, J. C. |author6=Fisher, K. |author7=Grigione, M. M. |author8=Lopez Gonzalez, C. A. |author9=Miller, J. R. |author10=Mormorunni, C. |author11=Paulson, L. |author12=Peters, R. |author13=Polisar, J. |author14=Povilitis, T. |author15=Robinson, M. J. |author16=Wilcox, S. |name-list-style=amp |title=The case for reintroduction: The jaguar (''Panthera onca'') in the United States as a model |journal=Conservation Science and Practice |date=2021 |volume=3 |issue=6 |page=e392 |doi=10.1111/csp2.392 |doi-access=free|bibcode=2021ConSP...3E.392S}}</ref>

==In culture and mythology==
{{multiple image |align=right |direction=vertical
|image1=Jaguar warrior.jpg |caption1=] in the Aztec culture
|image2=MocheJaguarLarcoMuseum.jpg |caption2=Moche jaguar figurine dating to 300&nbsp;], at the ] in ], Peru}}
{{Further|Jaguars in Mesoamerican cultures|Maya jaguar gods}}
In the ] Americas, the jaguar was a symbol of power and strength. In the Andes, a jaguar cult disseminated by the early ] became accepted over most of today's Peru by 900 ].<ref name="b1">{{cite book |title=The Earth and Its Peoples: A Global History |edition=Fifth |editor1=Bulliet, R.W. |editor2=Crossley, P. |editor3=Headrick, D. |editor4=Hirsch, S. |editor5=Johnson, L. |name-list-style=amp |year=2000 |publisher=Houghton Mifflin |volume=A 1: To 1200 |pages=75–76|url={{Google books|aujp0cT_TiEC|page=PA75|plainurl=yes}} |isbn=978-1-4390-8476-2}}</ref> The later ] in northern Peru used the jaguar as a symbol of power in many of their ceramics.<ref name="r41">{{cite book|author=Park, Yumi |year=2012 |title=Mirrors of Clay: Reflections of Ancient Andean Life in Ceramics from the Sam Olden Collection |publisher=]|page=49|isbn=9781617037955}}</ref> In the ] in ], the jaguar was considered a sacred animal, and people dressed in jaguar skins during religious rituals.<ref name="Ocampo_p231">{{cite book |last=Ocampo López |first=J. |author-link=Javier Ocampo López |year=2007 |title=Grandes culturas indígenas de América – Great indigenous cultures of the Americas |location=Bogotá, Colombia |publisher=Plaza & Janes Editores Colombia S.A. |language=es |page=231 |isbn=978-958-14-0368-4}}</ref>
The skins were traded with peoples in the nearby ].<ref>{{cite thesis |author=Kruschek, M.H. |year=2003 |type=PhD thesis |title=The evolution of the Bogotá chiefdom: A household view |publisher=] |location=Pittsburgh |url=http://d-scholarship.pitt.edu/7735/1/kruschek2003.pdf |access-date=12 January 2017 |archive-date=15 August 2016 |archive-url=https://web.archive.org/web/20160815211943/http://d-scholarship.pitt.edu/7735/1/kruschek2003.pdf |url-status=live }}</ref>
The name of the ] ] was derived from the ] words ''nymy'' and ''quyne'', meaning "force of the jaguar".<ref name="nymy">{{cite web |title=''nymy'' |publisher=Muysc cubun Dictionary Online |language=es |url=http://muysca.cubun.org/nymy |access-date=11 January 2017 |archive-date=13 October 2020 |archive-url=https://web.archive.org/web/20201013111835/http://muysca.cubun.org/nymy |url-status=live }}</ref><ref name="quyne">{{cite web |title=''quyne'' |publisher=Muysc cubun Dictionary Online |language=es |url=http://muysca.cubun.org/quyne |access-date=11 January 2017 |archive-date=10 January 2017 |archive-url=https://web.archive.org/web/20170110082526/http://muysca.cubun.org/quyne |url-status=live }}</ref>

Sculptures with "]" motifs were found on the Yucatán Peninsula in ] and ]; they show stylized jaguars with half-human faces.<ref>{{cite journal |author1=Metcalf, G. |name-list-style=amp |author2=Flannery, K.V. |year=1967 |title=An Olmec "were-jaguar" from the Yucatan Peninsula |journal=American Antiquity |volume=32 |issue=1 |pages=109–111 |doi=10.2307/278787 |jstor=278787|s2cid=164201137 }}</ref> In the later ], the jaguar was known as ''balam'' or ''bolom''' in many of the ], and was used to symbolize warriors and the elite class for being brave, fierce and strong.<ref name=Saunders/> The cat was associated with the ] and its image was used to decorate tombs and grave-good vessels.<ref name=Benson/>

The ] civilization called the jaguar ''ocelotl'' and considered it to be the king of the animals. It was believed to be fierce and courageous, but also wise, dignified and careful. The military had two classes of warriors, the ''ocelotl'' or ] and the ''cuauhtli'' or ]s and each dressed like their representative animal. In addition, members of the royal class would decorate in jaguar skins. The jaguar was considered to be the ] animal of the powerful deities ]<ref name=Saunders>{{cite journal |author=Saunders, N.J. |year=1994 |title=Predators of culture: Jaguar symbolism and Mesoamerican elites |journal=World Archaeology |volume=26 |issue=1 |pages=104–117 |doi=10.1080/00438243.1994.9980264 |jstor=124867}}</ref> and ].<ref name=Benson>{{cite book|last=Benson|first=Elizabeth P|contribution=The lord, the ruler: Jaguar symbolism in the Americas|year=2013|title=Icons of Power: Feline Symbolism in the Americas|editor=Saunders, Nicholas J.|pages=64–66|publisher=Routledge |isbn=9781136605130}}</ref>

A ] gorget depicting a jaguar was found in a ] in ]. The gorget shows evenly-engraved lines and measures {{cvt|104x98|mm}}.<ref name=Daggett/>
Rock drawings made by the ], ] and ] all over the desert and ] regions of the American Southwest show an explicitly spotted cat, presumably a jaguar, as it is drawn much larger than an ].<ref name=Pavlik /><!--commented out, as this ref is without date + publisher, hence gray literature: Jaguar skins were sold for $18 apiece in the mid 19th century in the vicinity of San Antonio, Texas.<ref>{{Cite report |author=Robinson, M. J. |title=Suitable Habitat for jaguars in New Mexico |publisher=location= |url=https://www.biologicaldiversity.org/species/mammals/jaguar/pdfs/NM_jaguar_habitat_report.pdf}}</ref>
-->

The jaguar is also used as a symbol in contemporary culture.<ref>{{Cite journal |last=Saunders |first=Nicholas J. |date=1994 |title=Predators of Culture: Jaguar Symbolism and Mesoamerican Elites |url=https://www.jstor.org/stable/124867 |journal=World Archaeology |volume=26 |issue=1 |pages=104–117 |doi=10.1080/00438243.1994.9980264 |jstor=124867 |issn=0043-8243 |access-date=29 January 2023 |archive-date=27 February 2023 |archive-url=https://web.archive.org/web/20230227075258/https://www.jstor.org/stable/124867 |url-status=live }}</ref> It is the ] of Guyana and is featured in its ].<ref>{{cite news |author=Khan, A. |date=2021 |title=National symbols: The Coat-Of-Arms |website=Guyana News and Information |url=http://www.guyana.org/Handbook/symbols.html |access-date=10 February 2021 |archive-date=12 May 2012 |archive-url=https://web.archive.org/web/20120512212328/http://guyana.org/Handbook/symbols.html |url-status=live }}</ref>
{{clear}}

==See also==
* ]

{{clear}}

== References ==
{{Reflist}}


==External links== ==External links==
{{commons| Panthera onca}} {{Commons category|Panthera onca}}
{{Wikispecies|Panthera onca}}
*ARKive -
{{Wiktionary|Panthera onca}}
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* {{cite web |title = Jaguar ''Panthera onca'' |publisher=IUCN Cat Specialist Group |url=http://www.catsg.org/index.php?id=95}}
* - Notecard
* {{cite web| title = Jaguars: Born free |work=BBC Natural World |year=2013 |url=https://www.bbc.co.uk/programmes/b01q9djl |access-date=13 August 2021}}
*
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* {{Cite Americana |short = x |wstitle = Jaguar}}

{{Spoken Misplaced Pages|Jaguar.ogg|date=26 December 2011}}
{{Carnivora|Fe.}}
{{Brazil symbols}}

{{Portal bar|Cats|Mammals|Animals|Biology|North America|Central America|South America|Mesoamerica}}

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Latest revision as of 13:56, 20 November 2024

Large cat native to the Americas For the car manufacturer, see Jaguar Cars. For other uses, see Jaguar (disambiguation).

Jaguar
Temporal range: Early Pleistocenepresent (~850,000–0 YBP)
Conservation status

Near Threatened  (IUCN 3.1)
CITES Appendix I (CITES)
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Feliformia
Family: Felidae
Subfamily: Pantherinae
Genus: Panthera
Species: P. onca
Binomial name
Panthera onca
(Linnaeus, 1758)
Subspecies
  Current range

  Former range

Synonyms
  • Felis augustus (Leidy, 1872)
  • Felis listai (Roth, 1899)
  • Felis onca Linnaeus, 1758
  • Felis onca subsp. boliviensis Nelson & Goldman, 1933
  • Felis onca subsp. coxi Nelson & Goldman, 1933
  • Felis onca subsp. ucayalae Nelson & Goldman, 1933
  • Felis veronis Hay, 1919
  • Iemish listai (Roth, 1899)
  • Panthera augusta (Leidy, 1872)
  • Panthera onca subsp. augusta (Leidy, 1872)
  • Uncia augusta (Leidy, 1872)

The jaguar (Panthera onca) is a large cat species and the only living member of the genus Panthera that is native to the Americas. With a body length of up to 1.85 m (6 ft 1 in) and a weight of up to 158 kg (348 lb), it is the biggest cat species in the Americas and the third largest in the world. Its distinctively marked coat features pale yellow to tan colored fur covered by spots that transition to rosettes on the sides, although a melanistic black coat appears in some individuals. The jaguar's powerful bite allows it to pierce the carapaces of turtles and tortoises, and to employ an unusual killing method: it bites directly through the skull of mammalian prey between the ears to deliver a fatal blow to the brain.

The modern jaguar's ancestors probably entered the Americas from Eurasia during the Early Pleistocene via the land bridge that once spanned the Bering Strait. Today, the jaguar's range extends from the Southwestern United States across Mexico and much of Central America, the Amazon rainforest and south to Paraguay and northern Argentina. It inhabits a variety of forested and open terrains, but its preferred habitat is tropical and subtropical moist broadleaf forest, wetlands and wooded regions. It is adept at swimming and is largely a solitary, opportunistic, stalk-and-ambush apex predator. As a keystone species, it plays an important role in stabilizing ecosystems and in regulating prey populations.

The jaguar is threatened by habitat loss, habitat fragmentation, poaching for trade with its body parts and killings in human–wildlife conflict situations, particularly with ranchers in Central and South America. It has been listed as Near Threatened on the IUCN Red List since 2002. The wild population is thought to have declined since the late 1990s. Priority areas for jaguar conservation comprise 51 Jaguar Conservation Units (JCUs), defined as large areas inhabited by at least 50 breeding jaguars. The JCUs are located in 36 geographic regions ranging from Mexico to Argentina.

The jaguar has featured prominently in the mythology of indigenous peoples of the Americas, including those of the Aztec and Maya civilizations.

Etymology

The word "jaguar" is possibly derived from the Tupi-Guarani word yaguara meaning 'wild beast that overcomes its prey at a bound'. In North America, the word is pronounced disyllabic /ˈdʒæɡwɑːr/, while in British English, it is pronounced with three syllables /ˈdʒæɡjuːər/. Because that word also applies to other animals, indigenous peoples in Guyana call it jaguareté, with the added sufix eté, meaning "true beast". "Onca" is derived from the Portuguese name onça for a spotted cat that is larger than a lynx; cf. ounce. The word "panther" is derived from classical Latin panthēra, itself from the ancient Greek πάνθηρ (pánthēr).

Taxonomy and evolution

Taxonomy

In 1758, Carl Linnaeus described the jaguar in his work Systema Naturae and gave it the scientific name Felis onca.

In the 19th and 20th centuries, several jaguar type specimens formed the basis for descriptions of subspecies. In 1939, Reginald Innes Pocock recognized eight subspecies based on the geographic origins and skull morphology of these specimens. Pocock did not have access to sufficient zoological specimens to critically evaluate their subspecific status but expressed doubt about the status of several. Later consideration of his work suggested only three subspecies should be recognized. The description of P. o. palustris was based on a fossil skull.

By 2005, nine subspecies were considered to be valid taxa:

Reginald Innes Pocock placed the jaguar in the genus Panthera and observed that it shares several morphological features with the leopard (P. pardus). He, therefore, concluded that they are most closely related to each other. Results of morphological and genetic research indicate a clinal north–south variation between populations, but no evidence for subspecific differentiation. DNA analysis of 84 jaguar samples from South America revealed that the gene flow between jaguar populations in Colombia was high in the past. Since 2017, the jaguar is considered to be a monotypic taxon, though the modern Panthera onca onca is still distinguished from two fossil subspecies, Panthera onca augusta and Panthera onca mesembrina. However, the 2024 study suggested that the validity of subspecific assignments on both P. o. augusta and P. o. mesembrina remains unresolved, since both fossil and living jaguars show a considerable variation in morphometry.

Evolution

Fossil skull of P. o. augusta

The Panthera lineage is estimated to have genetically diverged from the common ancestor of the Felidae around 9.32 to 4.47 million years ago to 11.75 to 0.97 million years ago. Some genetic analyzes place the jaguar as a sister species to the lion with which it diverged 3.46 to 1.22 million years ago, but other studies place the lion closer to the leopard.

The lineage of the jaguar appears to have originated in Africa and spread to Eurasia 1.95–1.77 mya. The living jaguar species is often suggested to have descended from the Eurasian Panthera gombaszoegensis. The ancestor of the jaguar entered the American continent via Beringia, the land bridge that once spanned the Bering Strait, Some authors have disputed the close relationship between P. gombaszoegensis (which is primarily known from Europe) and the modern jaguar. The oldest fossils of modern jaguars (P. onca) have been found in North America dating between 850,000-820,000 years ago. Results of mitochondrial DNA analysis of 37 jaguars indicate that current populations evolved between 510,000 and 280,000 years ago in northern South America and subsequently recolonized North and Central America after the extinction of jaguars there during the Late Pleistocene.

Two extinct subspecies of jaguar are recognized in the fossil record: the North American P. o. augusta and South American P. o. mesembrina.

Phylogenetic relationships of the jaguar as derived through analysis of
nuclear DNA:
Felidae

Felinae

Pantherinae
Panthera

Lion (P. leo)

Jaguar

Leopard (P. pardus)

Tiger (P. tigris)

Snow leopard (P. uncia)

Neofelis

mitochondrial DNA:
Felidae

Felinae

Pantherinae
Panthera

Lion

Leopard

Snow leopard

Jaguar

Tiger

Neofelis

Description

Skull of a jaguarIllustration of cheetah, leopard and jaguarA black jaguar. Such melanistic jaguars as well as leopards are commonly called black panthers

The jaguar is a compact and muscular animal. It is the largest cat native to the Americas and the third largest in the world, exceeded in size only by the tiger and the lion. It stands 57 to 81 cm (22.4 to 31.9 in) tall at the shoulders. Its size and weight vary considerably depending on sex and region: weights in most regions are normally in the range of 56–96 kg (123–212 lb). Exceptionally big males have been recorded to weigh as much as 158 kg (348 lb). The smallest females from Middle America weigh about 36 kg (79 lb). It is sexually dimorphic, with females typically being 10–20% smaller than males. The length from the nose to the base of the tail varies from 1.12 to 1.85 m (3 ft 8 in to 6 ft 1 in). The tail is 45 to 75 cm (18 to 30 in) long and the shortest of any big cat. Its muscular legs are shorter than the legs of other Panthera species with similar body weight.

Size tends to increase from north to south. Jaguars in the Chamela-Cuixmala Biosphere Reserve on the Pacific coast of central Mexico weighed around 50 kg (110 lb). Jaguars in Venezuela and Brazil are much larger, with average weights of about 95 kg (209 lb) in males and of about 56–78 kg (123–172 lb) in females.

The jaguar's coat ranges from pale yellow to tan or reddish-yellow, with a whitish underside and covered in black spots. The spots and their shapes vary: on the sides, they become rosettes which may include one or several dots. The spots on the head and neck are generally solid, as are those on the tail where they may merge to form bands near the end and create a black tip. They are elongated on the middle of the back, often connecting to create a median stripe, and blotchy on the belly. These patterns serve as camouflage in areas with dense vegetation and patchy shadows. Jaguars living in forests are often darker and considerably smaller than those living in open areas, possibly due to the smaller numbers of large, herbivorous prey in forest areas.

The jaguar closely resembles the leopard but is generally more robust, with stockier limbs and a more square head. The rosettes on a jaguar's coat are larger, darker, fewer in number and have thicker lines, with a small spot in the middle. It has powerful jaws with the third-highest bite force of all felids, after the tiger and the lion. It has an average bite force at the canine tip of 887.0 Newton and a bite force quotient at the canine tip of 118.6. A 100 kg (220 lb) jaguar can bite with a force of 4.939 kN (1,110 lbf) with the canine teeth and 6.922 kN (1,556 lbf) at the carnassial notch.

Color variation

Melanistic jaguars are also known as black panthers. The black morph is less common than the spotted one. Black jaguars have been documented in Central and South America. Melanism in the jaguar is caused by deletions in the melanocortin 1 receptor gene and inherited through a dominant allele. Black jaguars occur at higher densities in tropical rainforest and are more active during the daytime. This suggests that melanism provides camouflage in dense vegetation with high illumination.

In 2004, a camera trap in the Sierra Madre Occidental mountains photographed the first documented black jaguar in Northern Mexico. Black jaguars were also photographed in Costa Rica's Alberto Manuel Brenes Biological Reserve, in the mountains of the Cordillera de Talamanca, in Barbilla National Park and in eastern Panama.

Distribution and habitat

A female jaguar at Piquiri River, Mato Grosso state, BrazilA jaguar in São Lourenço River

In 1999, the jaguar's historic range at the turn of the 20th century was estimated at 19,000,000 km (7,300,000 sq mi), stretching from the southern United States through Central America to southern Argentina. By the turn of the 21st century, its global range had decreased to about 8,750,000 km (3,380,000 sq mi), with most declines occurring in the southern United States, northern Mexico, northern Brazil, and southern Argentina. Its present range extends from Mexico through Central America to South America comprising Belize, Guatemala, Honduras, Nicaragua, Costa Rica, particularly on the Osa Peninsula, Panama, Colombia, Venezuela, Guyana, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay and Argentina. It is considered to be locally extinct in El Salvador and Uruguay.

Jaguars have been occasionally sighted in Arizona, New Mexico and Texas, with 62 accounts reported in the 20th century. Between 2012 and 2015, a male vagrant jaguar was recorded in 23 locations in the Santa Rita Mountains. Eight jaguars were photographed in the southwestern US between 1996 and 2024.

The jaguar prefers dense forest and typically inhabits dry deciduous forests, tropical and subtropical moist broadleaf forests, rainforests and cloud forests in Central and South America; open, seasonally flooded wetlands, dry grassland and historically also oak forests in the United States. It has been recorded at elevations up to 3,800 m (12,500 ft) but avoids montane forests. It favors riverine habitat and swamps with dense vegetation cover. In the Mayan forests of Mexico and Guatemala, 11 GPS-collared jaguars preferred undisturbed dense habitat away from roads; females avoided even areas with low levels of human activity, whereas males appeared less disturbed by human population density. A young male jaguar was also recorded in the semi-arid Sierra de San Carlos at a waterhole.

Former range

In the 19th century, the jaguar was still sighted at the North Platte River 48–80 km (30–50 miles) north of Longs Peak in Colorado, in coastal Louisiana, northern Arizona and New Mexico. Multiple verified zoological reports of the jaguar are known in California, two as far north as Monterey in 1814 and 1826. The only record of an active jaguar den with breeding adults and kittens in the United States was in the Tehachapi Mountains of California prior to 1860. The jaguar persisted in California until about 1860. The last confirmed jaguar in Texas was shot in 1948, 4.8 km (3 miles) southeast of Kingsville, Texas. In Arizona, a female was shot in the White Mountains in 1963. By the late 1960s, the jaguar was thought to have been extirpated in the United States. Arizona outlawed jaguar hunting in 1969, but by then no females remained, and over the next 25 years only two males were sighted and killed in the state. In 1996, a rancher and hunting guide from Douglas, Arizona came across a jaguar in the Peloncillo Mountains and became a researcher on jaguars, placing trail cameras, which recorded four more jaguars.

Behavior and ecology

The jaguar is mostly active at night and during twilight. However, jaguars living in densely forested regions of the Amazon Rainforest and the Pantanal are largely active by day, whereas jaguars in the Atlantic Forest are primarily active by night. The activity pattern of the jaguar coincides with the activity of its main prey species. Jaguars are good swimmers and play and hunt in the water, possibly more than tigers. They have been recorded moving between islands and the shore. Jaguars are also good at climbing trees but do so less often than cougars.

Ecological role

Jaguar at Three Brothers River, Pantanal, Brazil

The adult jaguar is an apex predator, meaning it is at the top of the food chain and is not preyed upon in the wild. The jaguar has also been termed a keystone species, as it is assumed that it controls the population levels of prey such as herbivorous and seed-eating mammals and thus maintains the structural integrity of forest systems. However, field work has shown this may be natural variability, and the population increases may not be sustained. Thus, the keystone predator hypothesis is not accepted by all scientists.

The jaguar is sympatric with the cougar. In central Mexico, both prey on white-tailed deer, which makes up 54% and 66% of jaguar and cougar's prey, respectively. In northern Mexico, the jaguar and the cougar share the same habitat, and their diet overlaps dependent on prey availability. Jaguars seemed to prefer deer and calves. In Mexico and Central America, neither of the two cats are considered to be the dominant predator. In South America, the jaguar is larger than the cougar and tends to take larger prey, usually over 22 kg (49 lb). The cougar's prey usually weighs between 2 and 22 kg (4 and 49 lb), which is thought to be the reason for its smaller size. This situation may be advantageous to the cougar. Its broader prey niche, including its ability to take smaller prey, may give it an advantage over the jaguar in human-altered landscapes.

Hunting and diet

The jaguar has a powerful bite that allows it to pierce the shells of armored prey.
Jaguars killing and feeding on a yacare caiman

The jaguar is an obligate carnivore and depends solely on flesh for its nutrient requirements. An analysis of 53 studies documenting the diet of the jaguar revealed that its prey ranges in weight from 1 to 130 kg (2.2 to 286.6 lb); it prefers prey weighing 45–85 kg (99–187 lb), with the capybara and the giant anteater being the most selected. When available, it also preys on marsh deer, southern tamandua, collared peccary and black agouti. In floodplains, jaguars opportunistically take reptiles such as turtles and caimans. Consumption of reptiles appears to be more frequent in jaguars than in other big cats. One remote population in the Brazilian Pantanal is recorded to primarily feed on aquatic reptiles and fish. The jaguar also preys on livestock in cattle ranching areas where wild prey is scarce. The daily food requirement of a captive jaguar weighing 34 kg (75 lb) was estimated at 1.4 kg (3.1 lb) of meat.

The jaguar's bite force allows it to pierce the carapaces of the yellow-spotted Amazon river turtle and the yellow-footed tortoise. It employs an unusual killing method: it bites mammalian prey directly through the skull between the ears to deliver a fatal bite to the brain. It kills capybara by piercing its canine teeth through the temporal bones of its skull, breaking its zygomatic arch and mandible and penetrating its brain, often through the ears. It has been hypothesized to be an adaptation to cracking open turtle shells; armored reptiles may have formed an abundant prey base for the jaguar following the late Pleistocene extinctions. However, this is disputed, as even in areas where jaguars prey on reptiles, they are still taken relatively infrequently compared to mammals in spite of their greater abundance.

Between October 2001 and April 2004, 10 jaguars were monitored in the southern Pantanal. In the dry season from April to September, they killed prey at intervals ranging from one to seven days; and ranging from one to 16 days in the wet season from October to March.

The jaguar uses a stalk-and-ambush strategy when hunting rather than chasing prey. The cat will slowly walk down forest paths, listening for and stalking prey before rushing or ambushing. The jaguar attacks from cover and usually from a target's blind spot with a quick pounce; the species' ambushing abilities are considered nearly peerless in the animal kingdom by both indigenous people and field researchers and are probably a product of its role as an apex predator in several different environments. The ambush may include leaping into water after prey, as a jaguar is quite capable of carrying a large kill while swimming; its strength is such that carcasses as large as a heifer can be hauled up a tree to avoid flood levels. After killing prey, the jaguar will drag the carcass to a thicket or other secluded spot. It begins eating at the neck and chest. The heart and lungs are consumed, followed by the shoulders.

Social activity

Male (background) and young female (foreground) near the Cuiabá River, Porto Jofre, Poconé, Mato Grosso, Brazil

The jaguar is generally solitary except for females with cubs. In 1977, groups consisting of a male, female and cubs, and two females with two males were sighted several times in a study area in the Paraguay River valley. In some areas, males may form paired coalitions which together mark, defend and invade territories, find and mate with the same females and search for and share prey. A radio-collared female moved in a home range of 25–38 km (9.7–14.7 sq mi), which partly overlapped with another female. The home range of the male in this study area overlapped with several females.

The jaguar uses scrape marks, urine, and feces to mark its territory. The size of home ranges depends on the level of deforestation and human population density. The home ranges of females vary from 15.3 km (5.9 sq mi) in the Pantanal to 53.6 km (20.7 sq mi) in the Amazon to 233.5 km (90.2 sq mi) in the Atlantic Forest. Male jaguar home ranges vary from 25 km (9.7 sq mi) in the Pantanal to 180.3 km (69.6 sq mi) in the Amazon to 591.4 km (228.3 sq mi) in the Atlantic Forest and 807.4 km (311.7 sq mi) in the Cerrado. Studies employing GPS telemetry in 2003 and 2004 found densities of only six to seven jaguars per 100 km (62 mi) in the Pantanal region, compared with 10 to 11 using traditional methods; this suggests the widely used sampling methods may inflate the actual numbers of individuals in a sampling area. Fights between males occur but are rare, and avoidance behavior has been observed in the wild. In one wetland population with degraded territorial boundaries and more social proximity, adults of the same sex are more tolerant of each other and engage in more friendly and co-operative interactions.

Captive jaguar vocalizing while playing

The jaguar roars/grunts for long-distance communication; intensive bouts of counter-calling between individuals have been observed in the wild. This vocalization is described as "hoarse" with five or six guttural notes. Chuffing is produced by individuals when greeting, during courting, or by a mother comforting her cubs. This sound is described as low intensity snorts, possibly intended to signal tranquility and passivity. Cubs have been recorded bleating, gurgling and mewing.

Reproduction and life cycle

Jaguars mating in the northern Pantanal
Female jaguar picking up her cub

In captivity, the female jaguar is recorded to reach sexual maturity at the age of about 2.5 years. Estrus lasts 7–15 days with an estrus cycle of 41.8 to 52.6 days. During estrus, she exhibits increased restlessness with rolling and prolonged vocalizations. She is an induced ovulator but can also ovulate spontaneously. Gestation lasts 91 to 111 days. The male is sexually mature at the age of three to four years. His mean ejaculate volume is 8.6±1.3 ml. Generation length of the jaguar is 9.8 years.

In the Pantanal, breeding pairs were observed to stay together for up to five days. Females had one to two cubs. The young are born with closed eyes but open them after two weeks. Cubs are weaned at the age of three months but remain in the birth den for six months before leaving to accompany their mother on hunts. Jaguars remain with their mothers for up to two years. They appear to rarely live beyond 11 years, but captive individuals may live 22 years.

In 2001, a male jaguar killed and partially consumed two cubs in Emas National Park. DNA paternity testing of blood samples revealed that the male was the father of the cubs. Two more cases of infanticide were documented in the northern Pantanal in 2013. To defend against infanticide, the female may hide her cubs and distract the male with courtship behavior.

Attacks on humans

Further information: Man-eating animal § Jaguars

The Spanish conquistadors feared the jaguar. According to Charles Darwin, the indigenous peoples of South America stated that people did not need to fear the jaguar as long as capybaras were abundant. The first official record of a jaguar killing a human in Brazil dates to June 2008. Two children were attacked by jaguars in Guyana. The majority of known attacks on people happened when it had been cornered or wounded.

Threats

A South American jaguar killed by Theodore Roosevelt

The jaguar is threatened by loss and fragmentation of habitat, illegal killing in retaliation for livestock depredation and for illegal trade in jaguar body parts. It is listed as Near Threatened on the IUCN Red List since 2002, as the jaguar population has probably declined by 20–25% since the mid-1990s. Deforestation is a major threat to the jaguar across its range. Habitat loss was most rapid in drier regions such as the Argentine pampas, the arid grasslands of Mexico and the southwestern United States.

In 2002, it was estimated that the range of the jaguar had declined to about 46% of its range in the early 20th century. In 2018, it was estimated that its range had declined by 55% in the last century. The only remaining stronghold is the Amazon rainforest, a region that is rapidly being fragmented by deforestation. Between 2000 and 2012, forest loss in the jaguar range amounted to 83.759 km (32.340 sq mi), with fragmentation increasing in particular in corridors between Jaguar Conservation Units (JCUs). By 2014, direct linkages between two JCUs in Bolivia were lost, and two JCUs in northern Argentina became completely isolated due to deforestation.

In Mexico, the jaguar is primarily threatened by poaching. Its habitat is fragmented in northern Mexico, in the Gulf of Mexico and the Yucatán Peninsula, caused by changes in land use, construction of roads and tourism infrastructure. In Panama, 220 of 230 jaguars were killed in retaliation for predation on livestock between 1998 and 2014. In Venezuela, the jaguar was extirpated in about 26% of its range in the country since 1940, mostly in dry savannas and unproductive scrubland in the northeastern region of Anzoátegui. In Ecuador, the jaguar is threatened by reduced prey availability in areas where the expansion of the road network facilitated access of human hunters to forests. In the Alto Paraná Atlantic forests, at least 117 jaguars were killed in Iguaçu National Park and the adjacent Misiones Province between 1995 and 2008. Some Afro-Colombians in the Colombian Chocó Department hunt jaguars for consumption and sale of meat. Between 2008 and 2012, at least 15 jaguars were killed by livestock farmers in central Belize.

The international trade of jaguar skins boomed between the end of the Second World War and the early 1970s. Significant declines occurred in the 1960s, as more than 15,000 jaguars were yearly killed for their skins in the Brazilian Amazon alone; the trade in jaguar skins decreased since 1973 when the Convention on International Trade in Endangered Species was enacted. Interview surveys with 533 people in the northwestern Bolivian Amazon revealed that local people killed jaguars out of fear, in retaliation, and for trade. Between August 2016 and August 2019, jaguar skins and body parts were seen for sale in tourist markets in the Peruvian cities of Lima, Iquitos and Pucallpa. Human-wildlife conflict, opportunistic hunting and hunting for trade in domestic markets are key drivers for killing jaguars in Belize and Guatemala. Seizure reports indicate that at least 857 jaguars were involved in trade between 2012 and 2018, including 482 individuals in Bolivia alone; 31 jaguars were seized in China. Between 2014 and early 2019, 760 jaguar fangs were seized that originated in Bolivia and were destined for China. Undercover investigations revealed that the smuggling of jaguar body parts is run by Chinese residents in Bolivia.

Conservation

The jaguar is listed on CITES Appendix I, which means that all international commercial trade in jaguars or their body parts is prohibited. Hunting jaguars is prohibited in Argentina, Brazil, Colombia, French Guiana, Honduras, Nicaragua, Panama, Paraguay, Suriname, the United States, and Venezuela. Hunting jaguars is restricted in Guatemala and Peru. In Ecuador, hunting jaguars is prohibited, and it is classified as threatened with extinction. In Guyana, it is protected as an endangered species, and hunting it is illegal.

El Jefe, a jaguar in Arizona

In 1986, the Cockscomb Basin Wildlife Sanctuary was established in Belize as the world's first protected area for jaguar conservation.

Jaguar Conservation Units

In 1999, field scientists from 18 jaguar range countries determined the most important areas for long-term jaguar conservation based on the status of jaguar population units, stability of prey base and quality of habitat. These areas, called "Jaguar Conservation Units" (JCUs), are large enough for at least 50 breeding individuals and range in size from 566 to 67,598 km (219 to 26,100 sq mi); 51 JCUs were designated in 36 geographic regions including:

Optimal routes of travel between core jaguar population units were identified across its range in 2010 to implement wildlife corridors that connect JCUs. These corridors represent areas with the shortest distance between jaguar breeding populations, require the least possible energy input of dispersing individuals and pose a low mortality risk. They cover an area of 2,600,000 km (1,000,000 sq mi) and range in length from 3 to 1,102 km (1.9 to 684.8 mi) in Mexico and Central America and from 489.14 to 1,607 km (303.94 to 998.54 mi) in South America. Cooperation with local landowners and municipal, state, or federal agencies is essential to maintain connected populations and prevent fragmentation in both JCUs and corridors. Seven of 13 corridors in Mexico are functioning with a width of at least 14.25 km (8.85 mi) and a length of no more than 320 km (200 mi). The other corridors may hamper passage, as they are narrower and longer.

In August 2012, the United States Fish and Wildlife Service set aside 3,392.20 km (838,232 acres) in Arizona and New Mexico for the protection of the jaguar. The Jaguar Recovery Plan was published in April 2019, in which Interstate 10 is considered to form the northern boundary of the Jaguar Recovery Unit in Arizona and New Mexico.

In Mexico, a national conservation strategy was developed from 2005 on and published in 2016. The Mexican jaguar population increased from an estimated 4,000 individuals in 2010 to about 4,800 individuals in 2018. This increase is seen as a positive effect of conservation measures that were implemented in cooperation with governmental and non-governmental institutions and landowners.

An evaluation of JCUs from Mexico to Argentina revealed that they overlap with high-quality habitats of about 1,500 mammals to varying degrees. Since co-occurring mammals benefit from the JCU approach, the jaguar has been called an umbrella species. Central American JCUs overlap with the habitat of 187 of 304 regional endemic amphibian and reptile species, of which 19 amphibians occur only in the jaguar range.

Approaches

A jaguar in Belize

In setting up protected reserves, efforts generally also have to be focused on the surrounding areas, as jaguars are unlikely to confine themselves to the bounds of a reservation, especially if the population is increasing in size. Human attitudes in the areas surrounding reserves and laws and regulations to prevent poaching are essential to make conservation areas effective.

To estimate population sizes within specific areas and to keep track of individual jaguars, camera trapping and wildlife tracking telemetry are widely used, and feces are sought out with the help of detection dogs to study jaguar health and diet.

Current conservation efforts often focus on educating ranch owners and promoting ecotourism. Ecotourism setups are being used to generate public interest in charismatic animals such as the jaguar while at the same time generating revenue that can be used in conservation efforts. A key concern in jaguar ecotourism is the considerable habitat space the species requires. If ecotourism is used to aid in jaguar conservation, some considerations need to be made as to how existing ecosystems will be kept intact, or how new ecosystems will be put into place that are large enough to support a growing jaguar population.

Conservationists and professionals in Mexico and the United States have established the 56,000 acres (23,000 ha) Northern Jaguar Reserve in northern Mexico. Advocacy for reintroduction of the jaguar to its former range in Arizona and New Mexico have been supported by documentation of natural migrations by individual jaguars into the southern reaches of both states, the recency of extirpation from those regions by human action, and supportive arguments pertaining to biodiversity, ecological, human, and practical considerations.

In culture and mythology

Jaguar warrior in the Aztec cultureMoche jaguar figurine dating to 300 CE, at the Larco Museum in Lima, Peru Further information: Jaguars in Mesoamerican cultures and Maya jaguar gods

In the pre-Columbian Americas, the jaguar was a symbol of power and strength. In the Andes, a jaguar cult disseminated by the early Chavín culture became accepted over most of today's Peru by 900 BC. The later Moche culture in northern Peru used the jaguar as a symbol of power in many of their ceramics. In the Muisca religion in Altiplano Cundiboyacense, the jaguar was considered a sacred animal, and people dressed in jaguar skins during religious rituals. The skins were traded with peoples in the nearby Orinoquía Region. The name of the Muisca ruler Nemequene was derived from the Chibcha words nymy and quyne, meaning "force of the jaguar".

Sculptures with "Olmec were-jaguar" motifs were found on the Yucatán Peninsula in Veracruz and Tabasco; they show stylized jaguars with half-human faces. In the later Maya civilization, the jaguar was known as balam or bolom' in many of the Mayan languages, and was used to symbolize warriors and the elite class for being brave, fierce and strong. The cat was associated with the underworld and its image was used to decorate tombs and grave-good vessels.

The Aztec civilization called the jaguar ocelotl and considered it to be the king of the animals. It was believed to be fierce and courageous, but also wise, dignified and careful. The military had two classes of warriors, the ocelotl or jaguar warriors and the cuauhtli or eagle warriors and each dressed like their representative animal. In addition, members of the royal class would decorate in jaguar skins. The jaguar was considered to be the totem animal of the powerful deities Tezcatlipoca and Tepeyollotl.

A conch shell gorget depicting a jaguar was found in a burial mound in Benton County, Missouri. The gorget shows evenly-engraved lines and measures 104 mm × 98 mm (4.1 in × 3.9 in). Rock drawings made by the Hopi, Anasazi and Pueblo all over the desert and chaparral regions of the American Southwest show an explicitly spotted cat, presumably a jaguar, as it is drawn much larger than an ocelot.

The jaguar is also used as a symbol in contemporary culture. It is the national animal of Guyana and is featured in its coat of arms.

See also

References

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Extant Carnivora species
Feliformia ("cat-like" carnivorans)
Feliformia
Feloidea
Prionodon (Asiatic linsangs)
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Pantherinae
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