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Nyctereutes Temporal range: 5.332–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N Late Miocene - recent
Common and Japanese raccoon dog
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Canidae
Subfamily:	Caninae
Genus:	Nyctereutes Temminck, 1838
Type species
Canis viverrinus Temminck, 1838
Species
N. procyonoides N. viverrinus

Nyctereutes (Greek: nyx, nykt- "night" + ereutēs "wanderer") is a genus of canid which includes only two extant species, both known as raccoon dogs: the common raccoon dog (Nyctereutes procyonoides) and the Japanese raccoon dog (Nyctereutes viverrinus). Nyctereutes first entered the fossil record 5.5 million years ago (Mya) in northern China. It was one of the earliest canines to arrive in the Old World. All but two species became extinct before the end of the Pleistocene. A study suggests that the evolution of Nyctereutes was influenced by environmental and climatic changes, such as the expansion and contraction of forests and the fluctuations of temperature and precipitation.

Characteristics

They are typically recognized by their short snouts, round crania and the shaping of their molars, specifically the ratio between M1 and M2. Nyctereutes is considered mainly an opportunistic carnivore, feeding on small mammals, fish, birds, and insects, alongside occasional plants, specifically roots. Their diet is mostly influenced by environmental factors. Japanese raccoon dogs are considered distinct from the mainland species because of the larger skull size found in Russian and Hokkaido raccoon dogs.

Species

Extant species

Genus Nyctereutes – Temminck, 1838 – two species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Common raccoon dog	Nyctereutes procyonoides (Gray, 1834) Four subspecies N. p. procyonoides N. p. koreensis N. p. orestes N. p. ussuriensis	Mongolia, Russian Far East, China, Korea, Vietnam; introduced to Europe	Size: Habitat: Diet:	LC
Japanese raccoon dog	Nyctereutes viverrinus (Temminck, 1838)	Japan	Size: Habitat: Diet:	LC

Fossil species

• †Nyctereutes abdeslami 3.6—1.8 Mya (Morocco)
• †Nyctereutes donnezani 9.0—3.4 Mya (Eastern Europe, Spain)
• †Nyctereutes lockwoodi 3.42—3.2 Mya (Ethiopia)
• †Nyctereutes megamastoides (Europe)
• †Nyctereutes sinensis 3.6 Mya—781,000 years ago (Eastern Asia)
• †Nyctereutes tingi
• †Nyctereutes vinetorum

References

1. ^ Wozencraft, W. C. (2005). "Genus Nyctereutes". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 532–628. ISBN 978-0-8018-8221-0. OCLC 62265494.
2. Temminck, C.J. (1838). "Over de Kennis en de Verbreiding der Zoogdieren van Japan" [On the Knowledge and Dissemination of the Mammals of Japan]. Tijdschrift voor Natuurlijke Geschiedenis en Physiologie (in Dutch). 5: 273–293.
3. ^ Farjand, Arya; Zhang, Zhao-Qun; Liu, Wen-Hui; Jiao, Chen-Hui; Wang, Li-Hua (June 2021). "The evolution of Nyctereutes (Carnivora: Canidae) in the Nihewan Basin, Hebei, northern China". Palaeoworld. 30 (2): 373–381. doi:10.1016/j.palwor.2020.07.002. S2CID 225565692.
4. Kim, Sang-In; Oshida, Tatsuo; Lee, Hang; Min, Mi-Sook; Kimura, Junpei (2015). "Evolutionary and Biogeographical implications of variation in skull morphology of raccoon dogs (Nyctereutes procyonoides, Mammalia: Carnivora)". Biological Journal of the Linnean Society. 116 (1): 856–872. doi:10.1111/bij.12629.
5. Paleobiology Database: ''Nyctereutes abdeslami basic info.
6. Geraads, D.; Alemseged, Z.; et al. (2010). "Nyctereutes lockwoodi, n. sp., a new canid (Carnivora: Mammalia) from the middle Pliocene of Dikika, Lower Awash, Ethiopia". Journal of Vertebrate Paleontology. 30 (2): 981–987. doi:10.1080/02724631003758326. S2CID 83954055.
7. Paleobiology Database: ''Nyctereutes sinensis basic info.

Further reading

• Wayne, Robert K. (June 1993). "Molecular evolution of the dog family". Trends in Genetics. 9 (6): 218–224. doi:10.1016/0168-9525(93)90122-x. PMID 8337763.

v t e Extinct Canidae
Kingdom Animalia Phylum Chordata Class Mammalia Order: Carnivora Suborder: Caniformia
Canidae Canidae †Prohesperocyon †Hesperocyoninae Hesperocyon Mesocyon Cynodesmus Sunkahetanka Philotrox Enhydrocyon Paraenhydrocyon Caedocyon Ectopocynus Osbornodon O. brachypus O. fricki O. iamonensis O. renjiei O. sesnoni O. scitulus O. wangi †Borophaginae Archaeocyon Oxetocyon Otarocyon Rhizocyon Phlaocyonini Cynarctoides Phlaocyon P. achoros P. annectens P. latidens P. leucosteus P. mariae P. marslandensis P. minor P. multicuspus P. taylori P. yatkolai Borophagini Cormocyon Desmocyon Metatomarctus Euoplocyon Psalidocyon Microtomarctus Protomarctus Tephrocyon Cynarctina Paracynarctus Cynarctus Aelurodontina Tomarctus Aelurodon Borophagina Paratomarctus Carpocyon Protepicyon Epicyon Borophagus B. diversidens B. dudleyi B. hilli B. littoralis B. orc B. parvus B. pugnator B. secundus Caninae see below↓
Caninae Caninae †Leptocyon †Metalopex Urocyon †U. citrinus †U. galushai †U. minicephalus †U. progressus †U. webbi Vulpini †Ferrucyon †Prototocyon Nyctereutes (raccoon dogs) †N. abdeslami †N. donnezani †N. lockwoodi †N. megamastoides †N. sinensis †N. tingi †N. vinetorum Vulpes (true foxes) †V. hassani †V. praeglacialis †V. qiuzhudingi †V. riffautae †V. rooki †V. skinneri †V. stenognathus †V. gigas Canini (true dogs) †Canis gezi †Nurocyon Cerdocyonina (zorro) †Protocyon †Theriodictis Speothos †Pleistocene bush dog (S. pacivorus) †Dusicyon Falkland Islands wolf (D. australis) D. avus D. cultridens Lycalopex (South American foxes) †L. cultridens (?) †L. ensenadensis Canina (wolf-like canids) see below↓
Canina (wolf-like canids) Canina †Dire wolf (Aenocyon) †Sardinian dhole (Cynotherium) †Trinil dog (Mececyon) †Merriam's dog (Megacyon) †Xenocyon †Eucyon E. davisi E. debonisi E. ferox E. khoikhoi Lycaon †L. magnus †L. sekowei Dhole (Cuon alpinus) †C. a. antiquus †C. a. caucasicus †European dhole (C. a. europaeus) †Tien Shan dhole (C. a. hesperius) Canis †C. antonii †C. apolloniensis †Armbruster's wolf (C. armbrusteri) †C. arnensis †C. borjgali †C. cedazoensis †C. chihliensis †Edward's wolf (C. edwardii) †Etruscan wolf (C. etruscus) †C. lepophagus †Mosbach wolf (C. mosbachensis) †C. nehringi †C. palmidens †C. variabilis Coyote (C. latrans) †Pleistocene coyote (C. l. orcutti) Red wolf (C. rufus) †Florida black wolf (C. r. floridanus) †Gregory's wolf (C. r. gregoryi) Wolf (C. lupus) †Prehistoric C. l. bohemica C. l. lunellensis C. l. maximus C. l. santenaisiensis Cave wolf (C. l. spelaeus) Beringian wolf †Recently extinct Sicilian wolf (C. l. cristaldii) Hokkaido wolf (C. l. hattai) Japanese wolf (C. l. hodophilax) Austro-Hungarian wolf Synonymized under C. l. arctos: Bernard's wolf (C. l. bernardi) Synonymized under C. l. occidentalis: Kenai Peninsula wolf (C. l. alces) Manitoba wolf (C. l. griseoalbus) Synonymized under C. l. nubilus: Newfoundland wolf (C. l. beothucus) Cascade mountain wolf (C. l. fuscus) Mogollon mountain wolf (C. l. mogollonensis) Texas wolf (C. l. monstrabilis) Southern Rocky Mountain wolf (C. l. youngi)
See also: Evolution of the wolf Category:Extinct canids

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