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Huaxiagnathus

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Extinct genus of dinosaurs

Huaxiagnathus
Temporal range: Early Cretaceous, 125 Ma PreꞒ O S D C P T J K Pg N
Huaxiagnathus orientalis fossil displayed in Hong Kong Science Museum
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Family: Compsognathidae
Genus: Huaxiagnathus
Hwang et al., 2004
Species: H. orientalis
Binomial name
Huaxiagnathus orientalis
Hwang et al., 2004

Huaxiagnathus is a genus of theropod dinosaur from the Lower Cretaceous of China. It was a compsognathid, large for that group at about half a meter longer than Compsognathus and larger specimens of Sinosauropteryx, with the largest specimen about 1.8 meters (5.9 feet) in length.

The name Huaxiagnathus is derived from the Chinese Hua Xia, 華夏, a traditional word for "China", and from the Greek gnathos, Latinised into gnathus, meaning "jaw."

Description

Scale diagram of the two known specimens with a human (holotype in green)
Skeletal diagram

The holotype (CAGS-IG-02-301, Chinese Academy of Geological Sciences, Beijing) specimen was collected from the Yixian Formation (Jehol Group, Aptian) at Dabangou Village, Sihetun area, near Beipiao City, in western Liaoning Province. The holotype consists of an essentially complete skeleton, lacking only the end of the tail, preserved on five large slabs. Partially digested bones of an unidentified vertebrate were found within the holotype specimen. Another, larger specimen of Huaxiagnathus was discovered earlier in the Yixian Formation of the Sihetun area (NGMC 98-5-003, National Geological Museum of China, Beijing), but damage and mistakes made during its preparation rendered it unsuitable as a holotype.

Cladistic analysis indicates that Huaxiagnathus is the basal most known compsognathid, as indicated by its unspecialized forearm.

Hwang et al. (2004, pp. 14–15) diagnosed this genus as follows: differing from all other known compsognathids in having a very long posterior process of the premaxilla that overlaps the antorbital fossa, a manus equal to the combined lengths of the humerus and radius, large manual unguals I and II which are subequal in length and 167% the length of manual ungual III, a first metacarpal which has a smaller proximal transverse width than the second metacarpal, and the presence of a reduced olecranon process on the ulna. In 2024, Andrea Cau published a study on the phylogenetics of compsognathids and immature theropods in general that called the assessment of Huaxiagnathus within Compsognathidae into question. The paper recovered the taxon, along with four other proposed compsognathids in a polytomy within basal Coelurosauria. This polytomy notably did not include Composognathus proper, which would make none of these species compsognathids.

Phylogeny

Here is a simplified version of Cau (2024), which shows Huaxiagnathus in bold.

Tetanurae

Siamraptor

Siamotyrannus

Orionides

Streptospondylus

Xuanhanosaurus

Poekilopleuron

Piveteausaurus

Piatnitzkysaurus

Marshosaurus

Leshansaurus

Eustreptospondylus

Condorraptor

Asfaltovenator

Sciurumimus

Nedcolbertia

Magnosaurus

Duriavenator

Afrovenator

Compsognathus longipes

Compsognathus corallestris

Torvosaurus tanneri

Torvosaurus gurneyi

Megalosaurus

Scipionyx

Wiehenvenator

Iberospinus

Baryonychinae

Spinosaurinae

Avetheropoda

Allosauroidea (Incl. Juravenator at an indeterminate position)

Coelurosauria

Sinosauropteryx

Sinocalliopteryx

Mirischia

Huaxiagnathus

Zuolong

Tyrannoraptora

Maniraptoromorpha

NGMC 2124

Tanycolagreus

Stokesosaurus

Juratyrant

Eutyrannosauria

References

  1. ^ Hwang, S. H., Norell, M. A., Ji, Q., and Gao, K. (2004). "A large compsognathid from the Early Cretaceous Yixian Formation of China." Journal of Systematic Palaeontology, 2(1): 13-30.
  2. Cau, Andrea (2024). "A Unified Framework for Predatory Dinosaur Macroevolution" (PDF). Bollettino della Società Paleontologica Italiana. 63 (1): 1–19. doi:10.4435/BSPI.2024.08 (inactive 2024-11-20). Archived from the original (PDF) on 2024-04-27. Retrieved 2024-04-22.{{cite journal}}: CS1 maint: DOI inactive as of November 2024 (link)
Theropoda
Avemetatarsalia
Theropoda
    • see below↓
Theropoda
Theropoda
Neotheropoda
Coelophysoidea
Coelophysidae
Averostra
    • see below↓
Dubious neotheropods
Coelophysis bauri Dilophosaurus wetherilli
Averostra
Averostra
Ceratosauria
Ceratosauridae
Abelisauroidea
Noasauridae
Elaphrosaurinae
Noasaurinae
Abelisauridae
Majungasaurinae
Carnotaurinae
Brachyrostra
Furileusauria
Tetanurae
    • see below↓
Ceratosaurus nasicornis Limusaurus inextricabilis Rajasaurus narmadensis Aucasaurus garridoi
Tetanurae
Tetanurae
Orionides
Megalosauroidea
Piatnitzkysauridae
Megalosauria
Megalosauridae
Megalosaurinae
Afrovenatorinae
Spinosauridae
Baryonychinae
Ceratosuchopsini
Spinosaurinae
Spinosaurini
Avetheropoda
    • see below↓
Piatnitzkysaurus floresi

Torvosaurus tanneri

Spinosaurus aegyptiacus
Avetheropoda
Avetheropoda
Carnosauria
Allosauroidea
Metriacanthosauridae
Metriacanthosaurinae
Allosauria
Allosauridae
Carcharodontosauria
Neovenatoridae
Carcharodontosauridae
Carcharodontosaurinae
Giganotosaurini
Megaraptora?
Megaraptoridae
Coelurosauria
    • see below↓
Xuanhanosaurus qilixiaensis Allosaurus fragilis

Neovenator salerii Carcharodontosaurus saharicus

Australovenator wintonensis
Coelurosauria
Coelurosauria
Tyrannoraptora
Tyrannosauroidea
Coeluridae?
Proceratosauridae
Pantyrannosauria
Eutyrannosauria
Tyrannosauridae
Albertosaurinae
Tyrannosaurinae
Alioramini
Daspletosaurini
Teratophoneini
Tyrannosaurini
Maniraptoromorpha
    • see below↓
Dubious coelurosaurs
Zuolong salleei Stokesosaurus clevelandi

Alioramus remotus

Tarbosaurus bataar
Maniraptoromorpha
Maniraptoromorpha
Neocoelurosauria
Compsognathidae
Maniraptoriformes
Ornithomimosauria
Macrocheiriformes
Deinocheiridae
Ornithomimidae
Maniraptora
Sinosauropteryx prima

Deinocheirus mirificus

Qiupalong henanensis
Taxon identifiers
Huaxiagnathus
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