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Huaxiadraco

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(Redirected from Huaxiapterus benxiensis) Genus of tapejarid pterosaur

Huaxiadraco
Temporal range: Early Cretaceous, 120 Ma PreꞒ O S D C P T J K Pg N
Holotype specimen
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Suborder: Pterodactyloidea
Family: Tapejaridae
Subfamily: Sinopterinae
Genus: Huaxiadraco
Pêgas et al., 2023
Type species
Huaxiapterus corollatus
et al., 2006
Synonyms
  • Huaxiapterus corollatus
    (Lü et al., 2006)
  • Huaxiapterus benxiensis
    (Lü et al., 2007)

Huaxiadraco (meaning "Hua Xia dragon") is a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It is the third valid genus of tapejarid from the Jehol Biota, after Sinopterus and Eopteranodon. It contains one species, Huaxiadraco corollatus, originally assigned to the defunct genus Huaxiapterus.

History of discovery

Holotype of Huaxiapterus benxiensis, a junior synonym of H. corollatus

Huaxiadraco is based on the holotype ZMNH M8131, a nearly complete skeleton. It was originally assigned to the genus Huaxiapterus by Lü Junchang and colleagues in 2006, under the binomial name Huaxiapterus corollatus. Lü et al. also named another species, Huaxiapterus benxiensis, a year later. However, analyses have since found that these species are only distantly related to Huaxiapterus jii, the type species of Huaxiapterus, and thus require a new genus name. A 2023 review of Chinese tapejarids by Pêgas et al. have confirmed these analyses, finding most of them (including Huaxipterus jii) to belong to the coeval Sinopterus dongi, although corollatus was found to belong to a distinct genus. They thus created Huaxiadraco for corollatus, synonymized benxiensis with it, and referred specimens D2525 (previously considered Sinopterus), BMPC 103, 104, and 105 to it.

Classification

Based on their reassessment of the Sinopterus species-complex, Pêgas et al. modified their working dataset, previously used in the redescription of Aerotitan. Their cladogram is shown below:

Tapejaromorpha

Thalassodromidae

Tapejaridae
Caupedactylia

Caupedactylus ybaka

Aymberedactylus cearensis

Eutapejaria
Tapejarinae

Europejara olcadesorum

Tupandactylus imperator

Tupandactylus navigans

Tapejara wellnhoferi

Caiuajara dobruskii

Sinopterinae

Huaxiadraco corollatus

Eopteranodon lii

Sinopterus dongi

Wightia declivirostris

Afrotapejara zouhrii

Bakonydraco galaczi

References

  1. ^ Pêgas, R. V.; Zhoi, X.; Jin, X.; Wang, K.; Ma, W. (2023). "A taxonomic revision of the Sinopterus complex (Pterosauria, Tapejaridae) from the Early Cretaceous Jehol Biota, with the new genus Huaxiadraco". PeerJ. 11. e14829. doi:10.7717/peerj.14829. PMC 9922500.
  2. Lü, J.; Jin, X.; Unwin, D.M.; Zhao, L.; Azuma, Y.; Ji, Q. (2006). "A new species of Huaxiapterus (Pterosauria: Pterodactyloidea) from the Lower Cretaceous of western Liaoning, China with comments on the systematics of tapejarid pterosaurs". Acta Geologica Sinica. 80 (3): 315–326. doi:10.1111/j.1755-6724.2006.tb00251.x. S2CID 129851866.
  3. Junchang, Lü; Yubo, Gao; Lida, Xing; Zhixin, Li; Zhenyuan, Sun (2007). "New species of Huaxiapterus from the Early Cretaceous of western Liaoning". Acta Geologica Sinica - English Edition. 81 (5): 683–687. doi:10.1111/j.1755-6724.2007.tb00992.x. S2CID 128877390.
  4. Andres, B.; Clark, J.; Xu, X. (2014). "The Earliest Pterodactyloid and the Origin of the Group". Current Biology. 24 (9): 1011–6. doi:10.1016/j.cub.2014.03.030. PMID 24768054.
  5. Pinheiro, F.L.; Fortier, D.C.; Schultz, C.L.; De Andrade, J.A.F.G.; Bantim, R.A.M. (2011). "New information on Tupandactylus imperator, with comments on the relationships of Tapejaridae (Pterosauria)". Acta Palaeontologica Polonica. 56: 567–580. doi:10.4202/app.2010.0057.
  6. Pêgas, Rodrigo V.; Holgado, Borja; Ortiz David, Leonardo D.; Baiano, Mattia A.; Costa, Fabiana R. (January 2022). "On the pterosaur Aerotitan sudamericanus (Neuquén Basin, Upper Cretaceous of Argentina), with comments on azhdarchoid phylogeny and jaw anatomy". Cretaceous Research. 129: 104998. doi:10.1016/j.cretres.2021.104998. ISSN 0195-6671.
Pterosauria
Avemetatarsalia
Pterosauria
    • see below↓
Pterosauria
Pterosauria
Preondactylia
Caviramidae?
Austriadraconidae
Eopterosauria
Eudimorphodontoidea
Raeticodactylidae
Eudimorphodontidae
Eudimorphodontinae
Zambellisauria?
Macronychoptera
Dimorphodontidae
Lonchognatha?
Novialoidea
Campylognathoididae
Breviquartossa
Rhamphorhynchidae
Rhamphorhynchinae
Rhamphorhynchae
Angustinaripterini
Rhamphorhynchini
Digibrevisauria?
Scaphognathidae?
Pterodactylomorpha
    • see below↓
Campylognathoides liasicus

Scaphognathus crassirostris

Dorygnathus banthensis
Pterodactylomorpha
Pterodactylomorpha
Monofenestrata
Darwinoptera
Wukongopteridae
Wukongopterinae
Pterodactyliformes
Caelidracones
Anurognathidae
Anurognathinae
Batrachognathinae
Pterodactyloidea
Lophocratia
    • see below↓
Jeholopterus ninchengensis Kryptodrakon progenitor
Lophocratia
Archaeopterodactyloidea
Germanodactylidae
Euctenochasmatia
Ctenochasmatoidea
Gallodactylidae
Aurorazhdarchia
Aurorazhdarchidae
Ctenochasmatidae
Moganopterinae?
Gnathosaurinae
Ctenochasmatinae
Pterodaustrini
Eupterodactyloidea
Ornithocheiroidea
    • see below↓
Pterodactylus antiquus Plataleorhynchus streptorophorodon
Ornithocheiroidea
Ornithocheiroidea
Tapejaroidea
Dsungaripteridae
Dsungaripterinae
Azhdarchoidea
Tapejaromorpha
Thalassodromidae?
Tapejariformes
Caupedactylia?
Tapejaridae
Sinopterinae
Tapejarinae
Tapejarini
Caiuajarina
Neoazhdarchia
Dsungaripteromorpha?
Azhdarchomorpha
Neopterodactyloidea
Chaoyangopteridae
Chaoyangopterinae
Azhdarchiformes
Alanqidae?
Azhdarchidae
Azhdarchinae
Quetzalcoatlinae
Pteranodontoidea
    • see below↓
Bakonydraco galaczi

Tupandactylus imperator

Quetzalcoatlus
Pteranodontoidea
Pteranodontoidea
Pteranodontia
Pteranodontidae
Nyctosauromorpha
Aponyctosauria
Nyctosauridae
Ornithocheiromorpha
Lonchodectidae
Lanceodontia
Lonchodraconidae
Istiodactyliformes
Mimodactylidae
Istiodactylidae
Istiodactylinae
Ornithocheiriformes
Boreopteridae
Ornithocheirae
Ornithocheiridae
Ornithocheirinae
Targaryendraconia?
Cimoliopteridae
Targaryendraconidae
Anhangueria
Hamipteridae?
Anhangueridae
Coloborhynchinae?
Anhanguerinae
Tropeognathinae?
Tropeognathini
Mythungini
Pteranodon longiceps

Nyctosaurus gracilis

Ludodactylus sibbicki
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