Thyreocoridae - Misplaced Pages

Family of true bugs

Thyreocoridae

Alkindus atratus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hemiptera
Suborder:	Heteroptera
Infraorder:	Pentatomomorpha
Superfamily:	Pentatomoidea
Family:	Thyreocoridae Amyot & Serville, 1843
Subfamilies
Thyreocorinae Corimelaeninae

The Thyreocoridae are a family of shield bugs, known by common names that include negro bugs or ebony bugs. Historically, a few authors have called this family "Corimelaenidae" (e.g.), but the name Thyreocoridae, published in 1843, has nomenclatural priority over Corimelaenidae, published in 1872. Other classifications have placed them as a subfamily within the broad family Cydnidae.

Genera

There has been disagreement about how to treat subfamilies within the Thyreocoridae.

Alkindus Distant, 1889
Amyssonotum Horváth, 1919
Carrabas Distant, 1908
Corimelaena White, 1839
Cydnoides Malloch, 1919
Eumetopia Westwood, 1838
Galgupha Amyot & Serville, 1843
Godmania Horváth, 1919
Pericrepis Horváth, 1919
Pruhleria McAtee & Malloch, 1933
Strombosoma Amyot & Serville, 1843
Thyreocoris Schrank, 1801

References

"Thyreocoridae". Discover Life. Retrieved 2023-04-25.
Grazia, Jocelia; Schuh, Randall T.; Wheeler, Ward C. (2008). "Phylogenetic relationships of family groups in Pentatomoidea based on morphology and DNA sequences (Insecta: Heteroptera)" (PDF). Cladistics. 24 (6): 932–976. doi:10.1111/j.1096-0031.2008.00224.x. Archived (PDF) from the original on 10 January 2014.
Forero, Dimitri (2008). "The Systematics of the Hemiptera" (PDF). Revista Colombiana de Entomologia. 34 (1): 1–21, page 11. Archived (PDF) from the original on 10 January 2014.
Antônio R. Panizzi, Jocélia Grazia (2015) True Bugs (Heteroptera) of the Neotropics
Lis, Jerzy (2006) Thyreocoridae Amyot & Serville, 1843 – negro-bugs pp.148-149 In: Catalogue of the Heteroptera of the Palaearctic Region. Volume 5. Pentatomomorpha II, Chapter: Thyreocoridae Amyot & Serville, 1843 – negro-bugs, Publisher: The Netherlands Entomological Society, Wageningen, Editors: Aukema B., Rieger Ch. DOI: 10.13140/RG.2.1.3195.9127
David A. Rider, Cristiano Feldens Schwertner, Jitka Vilímová, Dávid Rédei, Petr Kment, Donald B. Thomas (2018) Higher Systematics of the Pentatomoidea. Chapter 2.2.18 in: Invasive Stink Bugs and Related Species (Pentatomoidea): Biology, Higher Systematics, Semiochemistry, and Management. J.E. McPherson, Ed. CRC Press DOI:10.1201/9781315371221-2
Dolling, W. R. (1981). "A rationalized classification of the burrower bugs (Cydnidae)". Systematic Entomology. 6 (1): 61–76. doi:10.1111/j.1365-3113.1981.tb00016.x.
Ahmad, Imtiaz & McPherson, J. E. (1990). "Male genitalia of the type species of Corimelaena White, Galgupha Amyot and Serville, and Cydnoides Malloch (Hemiptera: Cydnidae: Corimelaeninae) and their bearing on classification". Annals of the Entomological Society of America. 83 (2): 162–170.

Extant Hemiptera families

Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Subclass: Pterygota
Infraclass: Neoptera
Superorder: Paraneoptera

Suborder Auchenorrhyncha

Cicadomorpha

Cercopoidea (froghoppers)	Aphrophoridae Cercopidae Clastopteridae Epipygidae Machaerotidae
Cicadoidea (cicadas)	Cicadidae Tettigarctidae (hairy cicadas)
Membracoidea	Aetalionidae Cicadellidae (leafhoppers) Melizoderidae Membracidae (typical treehoppers, thorn bugs) Myerslopiidae

Fulgoromorpha
(planthoppers)

Fulgoroidea	Acanaloniidae Achilidae Achilixiidae Cixiidae Delphacidae Derbidae Dictyopharidae Eurybrachidae Flatidae Fulgoridae (lanternflies) Gengidae Hypochthonellidae Issidae Kinnaridae Lophopidae Meenoplidae Nogodinidae Ricaniidae Tettigometridae Tropiduchidae

Suborder Sternorrhyncha

Aleyrodoidea	Aleyrodidae (whiteflies)
Aphidoidea (aphids)	Aphididae
Coccoidea (scale insects)	Aclerdidae Asterolecaniidae (pit scales) Beesoniidae (beesoniids) Carayonemidae (carayonemids) Cerococcidae (ornate pit scales) Coccidae (soft scales) Conchaspididae Dactylopiidae (cochineals) Diaspididae (armored scales) Eriococcidae (felt scales) Halimococcidae (pupillarial palm scales) Kermesidae Kerriidae (lac scales) Lecanodiaspididae (false pit scales) Margarodidae (cottony cushion scales, giant coccids, ground pearls) Micrococcidae (Mediterranean scales) Monophlebidae (giant scales) Ortheziidae (ensign scales) Phenacoleachiidae (phenacoleachiids) Phoenicococcidae (palm scales) Pseudococcidae (mealybugs) Putoidae (giant mealybugs) Stictococcidae (stictococcids)
Phylloxeroidea	Adelgidae (woolly conifer aphids) Phylloxeridae (phylloxerans)
Psylloidea	Aphalaridae Calophyidae Carsidaridae Homotomidae Phacopteronidae Psyllidae (jumping plant lice) Triozidae

Suborder Heteroptera (with Coleorrhyncha)

Dipsocoromorpha	Ceratocombidae Dipsocoridae Hypsipterygidae Schizopteridae Stemmocryptidae

Enicocephalomorpha

Enicocephaloidea	Aenictopecheidae Enicocephalidae (unique-headed bugs, gnat bugs)

Gerromorpha
(semiaquatic bugs)

Gerroidea	Gerridae (water striders) Hermatobatidae Veliidae (riffle bugs)
Hebroidea	Hebridae (velvet water bugs)
Hydrometroidea	Hydrometridae (marsh treaders or water measurers) Macroveliidae Paraphrynoveliidae
Mesovelioidea (water treaders)	Madeoveliidae Mesoveliidae

Leptopodomorpha	Saldidae (shore bugs) Leptopodidae (spiny shore bugs) Omaniidae Aepophilidae

Nepomorpha
(true water bugs)

Corixoidea	Corixidae (water boatmen)
Nepoidea	Belostomatidae (giant water bugs) Nepidae (water scorpions, needle bugs)
Ochteroidea	Gelastocoridae (toad bugs) Ochteridae (velvety shore bugs)
Aphelocheiroidea	Aphelocheiridae Potamocoridae
Naucoroidea	Naucoridae (creeping water bugs)
Notonectoidea	Notonectidae (backswimmers)
Pleoidea	Helotrephidae Pleidae (pygmy backswimmers)

Peloridiomorpha (Coleorrhyncha)	Peloridiidae

Cimicomorpha

Cimicoidea

Pentatomomorpha

Aradoidea	Aradidae (flat bugs) Termitaphididae (termite bugs)
Pentatomoidea (shield bugs)	Urostylididae Acanthosomatidae (shield bugs) Tessaratomidae (giant shield bugs and relatives) Dinidoridae Cydnidae (burrowing bugs) Thaumastellidae Parastrachiidae Thyreocoridae (includes ebony bugs) Lestoniidae Phloeidae Scutelleridae (jewel bugs or metallic shield bugs) Plataspidae Pentatomidae (stink bugs) Canopidae Megarididae
Coreoidea	Alydidae (broad-headed bugs) Coreidae (squash bugs, leaf-footed bugs) Hyocephalidae Rhopalidae (scentless plant bugs) Stenocephalidae
Lygaeoidea	Artheneidae Berytidae (stilt bugs) Blissidae Colobathristidae Cryptorhamphidae Cymidae Geocoridae Henicocoridae Heterogastridae Idiostolidae Lygaeidae (milkweed bugs, true seed bugs) Malcidae Ninidae Oxycarenidae Pachygronthidae Rhyparochromidae (atypical seed bugs) Piesmatidae (ash-grey leaf bugs)
Pyrrhocoroidea	Pyrrhocoridae (red bugs, cotton stainers) Largidae (bordered plant bugs)

Note: Coleorrhyncha are a different clade from Heteroptera. Heteroptera with Coleorrhyncha were referred to as Prosorrhyncha.

Taxon identifiers
Thyreocoridae	Wikidata: Q7799676 Wikispecies: Thyreocoridae BOLD: 198302 BugGuide: 6985 CoL: H8R EoL: 641 Fauna Europaea: 12854 Fauna Europaea (new): e58d9a11-c006-41bc-9dbf-c459879888d5 GBIF: 7898 iNaturalist: 133023 IRMNG: 111236 ITIS: 109110 NBN: NHMSYS0020309892 NCBI: 468275 Open Tree of Life: 658170 Plazi: 20224328-620E-032E-BC0A-FF7AFC78ADCA

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