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Revision as of 05:41, 21 April 2022

Extinct superfamily of dinosaurs

Coelophysoids
Temporal range: Late Triassic-Early Jurassic, 221–183 Ma PreꞒ O S D C P T J K Pg N
Mounted skeleton of Coelophysis bauri, Cleveland Museum of Natural History
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Neotheropoda
Superfamily: Coelophysoidea
Nopcsa, 1928
Type species
Coelurus bauri
Cope, 1887
Subgroups
Synonyms
  • Podokesauroidea Huene, 1914

Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the coelurosaurs, with which they were formerly classified, and some species had delicate cranial crests. Sizes range from about 1 to 6 m in length. It is unknown what kind of external covering coelophysoids had, and various artists have portrayed them as either scaly or feathered. Some species may have lived in packs, as inferred from sites where numerous individuals have been found together.

Examples of coelophysoids include Coelophysis, Procompsognathus and Liliensternus. Most dinosaurs formerly referred to as being in the dubious taxon "Podokesauridae" are now classified as coelophysoids.

Classification

Despite their very early occurrence in the fossil record, coelophysoids have a number of derived features that separate them from primitive (basal) theropods. Among the most prominent of these derived features (apomorphies) is the way the upper jaw bones are connected (the premaxilla-maxilla articulation), which is flexible with a deep gap between the teeth in the two bones. A major source of disagreement among theropod experts is whether or not coelophysoids shared a more recent common ancestor with Ceratosauria (sensu stricto) than the ceratosaurs did with other theropods. Most recent analyses indicate the latter, that Coelophysoidea does not form a natural group with the ceratosaurians. Similarly, while Dilophosaurus and similar theropods have traditionally been classified as coelophysoids, several studies published in the late 2000s suggested that they may actually be more closely related to the tetanurans.

The cladogram below was recovered in a study by Matthew T. Carrano, John R. Hutchinson and Scott D. Sampson, 2005.

Coelophysoidea

Dilophosaurus

Sarcosaurus

Gojirasaurus

Zupaysaurus

Coelophysidae

Procompsognathus

Segisaurus

Liliensternus

Coelophysis

Megapnosaurus

The cladogram below was recovered in a study by Martin D. Ezcurra and Gilles Cuny, 2007.

Liliensternus

Lophostropheus

 Coelophysidae 

"Megapnosaurus" kayentakatae

Coelophysis

Coelophysis rhodesiensis

The cladogram below was recovered in a study by Ezcurra et al. (2020).

Coelophysoidea

Liliensternus

Dracoraptor

'Syntarsus' kayentakatae

Panguraptor

Powellvenator

Lepidus

Coelophysis bauri

Camposaurus

Lucianovenator

Megapnosaurus rhodesiensis

Segisaurus

See also

References

  1. Smith, N.D., Makovicky, P.J., Pol, D., Hammer, W.R., and Currie, P.J. (2007). "The dinosaurs of the Early Jurassic Hanson Formation of the Central Transantarctic Mountains: Phylogenetic review and synthesis." In Cooper, A.K. and Raymond, C.R. et al. (eds.), Antarctica: A Keystone in a Changing World––Online Proceedings of the 10th ISAES, USGS Open-File Report 2007-1047, Short Research Paper 003, 5 p.; doi:10.3133/of2007-1047.srp003.
  2. Carrano, M.T, Hutchinson, J.R, Sampson, S.D. (2005). "New information on Segisaurus halli, a small theropod dinosaur from the Early Jurassic of Arizona." Journal of Vertebrate Paleontology. 25(4):835-849.
  3. Ezcurra, Martin D.; Cuny, Gilles (2007). "The coelophysoid Lophostropheus airelensis, gen. nov.: a review of the systematics of "Liliensternus" airelensis from the Triassic-Jurassic boundary outcrops of Normandy (France)". Journal of Vertebrate Paleontology. 27 (1): 73–86. doi:10.1671/0272-4634(2007)27[73:TCLAGN]2.0.CO;2.
  4. Ezcurra, Martín D; Butler, Richard J; Maidment, Susannah C R; Sansom, Ivan J; Meade, Luke E; Radley, Jonathan D (2021-01-01). "A revision of the early neotheropod genus Sarcosaurus from the Early Jurassic (Hettangian–Sinemurian) of central England". Zoological Journal of the Linnean Society. 191 (1): 113–149. doi:10.1093/zoolinnean/zlaa054. ISSN 0024-4082.

Sources

  • Rauhut and Hungerbuhler (2000). "A review of European Triassic theropods." Gaia, 15: 75-88.
  • Tykoski, R. S. (2005). "Anatomy, Ontogeny, and Phylogeny of Coelophysoid Theropods." Ph. D dissertation.
  • Yates, A.M., 2006 (for 2005). "A new theropod dinosaur from the Early Jurassic of South Africa and its implications for the early evolution of theropods." Palaeontologia Africana, 41: 105-122.
Theropoda
Avemetatarsalia
Theropoda
    • see below↓
Theropoda
Theropoda
Neotheropoda
Coelophysoidea
Coelophysidae
Averostra
    • see below↓
Dubious neotheropods
Coelophysis bauri Dilophosaurus wetherilli
Averostra
Averostra
Ceratosauria
Ceratosauridae
Abelisauroidea
Noasauridae
Elaphrosaurinae
Noasaurinae
Abelisauridae
Majungasaurinae
Carnotaurinae
Brachyrostra
Furileusauria
Tetanurae
    • see below↓
Ceratosaurus nasicornis Limusaurus inextricabilis Rajasaurus narmadensis Aucasaurus garridoi
Tetanurae
Tetanurae
Orionides
Megalosauroidea
Piatnitzkysauridae
Megalosauria
Megalosauridae
Megalosaurinae
Afrovenatorinae
Spinosauridae
Baryonychinae
Ceratosuchopsini
Spinosaurinae
Spinosaurini
Avetheropoda
    • see below↓
Piatnitzkysaurus floresi

Torvosaurus tanneri

Spinosaurus aegyptiacus
Avetheropoda
Avetheropoda
Carnosauria
Allosauroidea
Metriacanthosauridae
Metriacanthosaurinae
Allosauria
Allosauridae
Carcharodontosauria
Neovenatoridae
Carcharodontosauridae
Carcharodontosaurinae
Giganotosaurini
Megaraptora?
Megaraptoridae
Coelurosauria
    • see below↓
Xuanhanosaurus qilixiaensis Allosaurus fragilis

Neovenator salerii Carcharodontosaurus saharicus

Australovenator wintonensis
Coelurosauria
Coelurosauria
Tyrannoraptora
Tyrannosauroidea
Coeluridae?
Proceratosauridae
Pantyrannosauria
Eutyrannosauria
Tyrannosauridae
Albertosaurinae
Tyrannosaurinae
Alioramini
Daspletosaurini
Teratophoneini
Tyrannosaurini
Maniraptoromorpha
    • see below↓
Dubious coelurosaurs
Zuolong salleei Stokesosaurus clevelandi

Alioramus remotus

Tarbosaurus bataar
Maniraptoromorpha
Maniraptoromorpha
Neocoelurosauria
Compsognathidae
Maniraptoriformes
Ornithomimosauria
Macrocheiriformes
Deinocheiridae
Ornithomimidae
Maniraptora
Sinosauropteryx prima

Deinocheirus mirificus

Qiupalong henanensis
Taxon identifiers
Coelophysoidea
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