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Sphenocoelus Temporal range: Middle Eocene
Fossil
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Perissodactyla
Family:	†Brontotheriidae
Genus:	†Sphenocoelus Osborn, 1895
Species
See text

Sphenocoelus is an extinct genus of brontothere endemic to North America during the Middle Eocene 46.2—40.4 mya), existing for approximately 6 million years. Fossils have been found only in southern Wyoming and eastern Utah.

In life, it would have resembled a hornless rhinoceros, to which it was distantly related. However, it was smaller than modern rhinos, standing about 1.2 metres (3.9 ft) high at the shoulders, and had a much longer head. Its teeth were adapted to eating soft vegetation, such as forest leaves. Like other brontotheres, it had four toes on the front feet, and three on the hind feet, and the animal is unlikely to have been able to run fast.

Species

• S. blairi
• S. bridgeri
• S. harundivoras
• S. hyognathus (syn. Dolichorhinus longiceps, Telmatotherium cornutum)
• S. intermedius (syn. Dolichorhinus fluminalis, D. heterodon, Mesatirhinus superior)
• S. uintensis.

References

1. Sphenocoelus at fossilworks
2. Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. XXX. ISBN 1-84028-152-9.

v t e Perissodactyla
Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Perissodactyla
Perissodactyla Incertae sedis †Ghazijhippus †Mesolambdolophus †Olbitherium †Anthracobunia Radinskya †Anthracobunidae Anthracobune Hsanotherium Jozaria Obergfellia †Cambaytheriidae Cambaytherium Kalitherium Nakusia Perissobune †Brontotheriidae See Brontotheriidae Equoidea †Cymbalophus †Hallensia †Pliolophus †Systemodon Equidae See Equidae †Palaeotheriidae †Pachynolophinae Anchilophus Bepitherium Eurohippus Hyracotherium Lijiangia Lophiohippus Lophiotherium Metanchilophus Orolophus Pachynolophus Paranchilophus Propalaeotherium Qianohippus †Palaeotheriinae Cantabrotherium Iberolophus Idiodontherium Leptolophus Mekodontherium Metaplagiolophus Palaeotherium Paraplagiolophus Plagiolophus Pseudopalaeotherium †Hyopsodontidae? Aletodon Chacomylus Dorraletes Haplaletes Haplomylus Hyopsodus Litomylus Louisina Monshyus Paratricuspiodon Paschatherium Phenacodaptes Tricuspiodon †Phenacodontidae? Tetraclaenodon †Meniscotheriinae Ectocion Meniscotherium †Phenacodontinae Almogaver Copecion Eodesmatodon Lophocion Phenacodus Tapiromorpha See below↓
Tapiromorpha Incertae sedis †Cambaylophus †Skopaiolophus †Isectolophidae Chowliia Gandheralophus Karagalax Meridiolophus Orientolophus Sastrilophus †Homogalaxinae Homogalax †Isectolophinae Isectolophus Schizolophodon †Ancylopoda See Ancylopoda Ceratomorpha Tapiroidea †Deperetellidae Bahinolophus Deperetella Irenolophus Teleolophus †Helaletidae Colodon Dilophodon Helaletes Heptodon Heteraletes Paracolodon Plesiocolopirus Thuliadanta? Vastanolophus Tapiridae †Eotapirus †Miotapirus †Nexuotapirus †Paratapirus †Plesiotapirus †Protapirus †Tapiravus Tapirus Rhinocerotoidea See below↓
Rhinocerotoidea Rhinocerotoidea sensu lato †Breviodon †Fouchia †Minchenoletes †Siamolophus †Triplopus †Yimengia †Indolophidae Indolophus Rhinocerotoidea sensu stricto †Selenaletes †Hyrachyidae Hyrachyus †Eggysodontidae Allacerops Eggysodon Guangnanodon Proeggysodon †Hyracodontidae Ardynia Ephyrachyus Epitriplophus Hyracodon Prohyracodon Rhodopagus Triplopides †Amynodontidae Amynodontopsis Armania Cadurcotherium Caenolophus Huananodon Hypsamynodon Procadurcodon Rostriamynodon Teilhardia †Amynodontinae Amynodon †Cadurcodontini Cadurcodon Lushiamynodon Sharamynodon Sianodon †Metamynodontini Gigantamynodon Megalamynodon Metamynodon Paramynodon Zaisanamynodon †Paraceratheriidae †Forstercooperiinae Forstercooperia Gobicerops Pappaceras †Paraceratheriinae Aralotherium Dzungarotherium Juxia Paraceratherium Urtinotherium Rhinocerotidae See below↓
Rhinocerotidae Rhinocerotidae †Amphicaenopus †Epiaceratherium †Guixia †Teletaceras †Uintaceras †Woodoceras †Aceratheriinae Aceratherium Acerorhinus Alicornops Aphelops Chilotheridium Chilotherium Floridaceras Galushaceras Hoploaceratherium Mesaceratherium Peraceras Persiatherium Plesiaceratherium Ronzotherium Shansirhinus Sinorhinus Subchilotherium †Elasmotheriinae Gulfoceras Victoriaceros †Diceratheriini Diceratherium Penetrigonias Skinneroceras Subhyracodon Trigonias †Elasmotheriini Bugtirhinus Caementodon Elasmotherium Hispanotherium Iranotherium Kenyatherium Meninatherium Menoceras Ninxiatherium Ningxiatherium Ougandatherium Parelasmotherium Procoelodonta Sinotherium Tongxinotherium Rhinocerotinae †Dromoceratherium †Lartetotherium †Molassitherium †Protaceratherium †Teleoceratini Aprotodon Brachydiceratherium Brachypotherium Diaceratherium Prosantorhinus Shennongtherium Teleoceras Rhinocerotina Dicerorhinini †Coelodonta Dicerorhinus †Dihoplus †Pliorhinus †Stephanorhinus Dicerotini Ceratotherium Diceros †Miodiceros †Paradiceros Rhinocerotini †Gaindatherium †Nesorhinus Rhinoceros †Rusingaceros

v t e Brontotheriidae
Kingdom: Animalia Phylum: Chordata Clade: Synapsida Class: Mammalia Order: Perissodactyla Suborder: Hippomorpha
Basal brontotheres	Danjiangia Eotitanops Lambdotherium Mulkrajanops Pakotitanops Palaeosyops
Brontotheriinae	Acrotitan Arctotitan Bunobrontops Desmatotitan Dolichorhinus Fossendorhinus Hyotitan Mesatirhinus Metarhinus Metatelmatherium Microtitan Nanotitanops Pygmaetitan Qufutitan Sphenocoelus Sthenodectes Telmatherium Brontotheriini Diplacodon Protitan Protitanotherium Rhinotitan Pachytitan Brachydiastematherium Sivatitanops Embolotheriina Aktautitan Embolotherium Gnathotitan Maobrontops Metatitan Nasamplus Protembolotherium Brontotheriina Diantotitan Duchesneodus Epimanteoceras Eubrontotherium Megacerops Notiotitanops Parabrontops Parvicornus Protitanops

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